Rutaceae

Citrus japonica Thunb. forma spinosa Miq., Ann. Mus. Lugd.-Bat. 2: 83 (1865); Prolus. Fl. Jap.: 15 (1865).
Present treatment. Fortunella japonica (Thunb.) Swingle.
Type material. In m.[monte] Kawara Jama ins. Kiusiu, Pierot [no.103] [908. 203-185], holotype, with Tyozaburo Tanaka's determination as “Fortunella japonica (Thunb.) Swingle” in 1927.
Dictamnus fraxinella Pers. [var.] α parvifolia Miq., Ann. Mus. Lugd.-Bat. 3: 22 (1867); Prolus. Fl. Jap.: 210 (1867).
Present treatment. Dictamnus fraxinella Pers.
Type material. Siebold s.n. [908. 202-359], holotype.
Dictamnus fraxinella Pers. [var.] β oblongifolia Miq., Ann. Mus. Lugd.-Bat. 3: 22 (1867); Prolus. Fl. Jap.: 210 (1867).
Present treatment. Dictamnus fraxinella Pers.
Type material. Ex Archipelago Coreano, Oldham no. 126 [908. 203-914], holotype.
Dictamnus fraxinella Pers. [var.] γ pubescens Miq., Ann. Mus. Lugd.-Bat. 3: 22 (1867); Prolus. Fl. Jap.: 210 (1867).
Present treatment. Dictamnus fraxinella Pers.
Type material. Siebold s.n. [908. 203-915] (L 0367032), holotype.
Euodia glauca Miq., Ann. Mus. Lugd.-Bat 3: 23 (1867); Prolus. Fl. Jap.: 211 (1867). Hara, Enum. Sperm. Jap. 3: 24 (1954).
Present treatment. Tetradium glabrifolium (Benth.) T. G. Hartley var. glaucum (Miq.) H. Ohba: Ohba in Fl. Jap. 2c: 35 (1999).
Type materials. Buerger s.n. [908. 203-276] (L 0017996), lectotype (here chosen); [908. 203-275] (L 0017993), syntype; [908. 203-279] (L 0017994), syntype; [908. 203-285] (L 0017990), syntype; [908. 203-293] (L 0017995), syntype; [908. 203-300] (L 0017989), syntype. Detexit botanicus indigenus, probabiliter medicus Jedoensis, utrumque sexum, syntype [not found]. Mohnike s.n. [908. 203-263] (L 0017986), syntype. Pierot s.n. [908. 203-289] (L 0017997), excl. the middle branch (this is Phellodendron), syntype. Prope Nangasaki & probabiliter in Archipelago Coreano, Oldham no. 130, syntype [not found].
Other material. Unknown coll. s.n. [without registered no.] (not included in the database).
Ruta subtripinnata Miq., Ann. Mus. Lugd.-Bat. 3: 21 (1867); Prolus. Fl. Jap.: 209 (1867).
Present treatment. Ruta chalepensis L.
Type materials. Buerger s.n. [908. 203-1658] (L 0421294), lectotype (here chosen); [908. 203-1678] (L 0421293), isolectotype. In Japonia introducta, Siebold s.n. [908. 203-1601] (L 0421291), syntype; [908. 203-1676] (L 0421295), syntype; [908. 203-1681] (L 0421288), syntype. In ripa[ripais] fluvii Rokfgo Gawa insulae Nippon, Pierot [no. 27] [908. 203-1659] (L 0421289), syntype. Mohnike s.n. [908. 203-1657] (L 0421290), syntype; [908. 203-1679] (L 0421292).
Other materials. Keiske s.n. [908. 203-1677] (L 0175847). Pierot s.n. [908. 203-1660] (L 0421296).
Zanthoxylum emarginellum Miq., Ann. Mus. Lugd.-Bat. 3: 22 (1867); Prolus. Fl. Jap.: 210 (1867).
Fagara ailanthoides (Siebold & Zucc.) Engl.: Hara, Enum. Sperm. Jap. 3: 25 (1954).
Present treatment. Zanthoxylum ailanthoides Siebold & Zucc.(var. ailanthoides): Ohba in Fl. Jap. 2c: 39 (1999).
Type material. Siebold s.n. (from a Japanese botanist) [908. 204-113] (L 0424424), holotype.

Meliaceae

Melia azedarach L. var. subtripinnata Miq., Ann. Mus. Lugd.-Bat. 3: 24 (1867); Prolus. Fl. Jap.: 212 (1867). Hara, Enum. Sperm. Jap. 3: 36 (1954).
Present treatment. Melia azedarach L.: Iwatsuki & Ohba in Fl. Jap. 2c: 49 (1999).
Type materials. Japan. Keiske s.n. [908. 133-729] (L 0042886), lectotype (here chosen). Buerger, syntype [not found].
Other material. Siebold s.n. [908. 133-741] (L 0103833).

Polygalaceae

Polygala* sieboldiana Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 85 (1866); Ann. Mus. Lugd.-Bat. 2: 260 (1866); Prolus. Fl. Jap.: 148 (1866).
Polygala japonica Houtt. var. angustifolia Koidz., Fl. Symb. Orient.-Asiat.: 8 (1930).
Present treatment. Polygala tatarinowii Regel: Hara, Enum. Sperm. Jap. 3: 38 (1954). Iwatsuki & Ohba in Fl. Jap. 2c: 54 (1999).
Type material. Botanicus japonensis cum Sieboldo communicavit, Siebold s.n. [908. 171-1884] (L 0017884), with F. Adema's determination as P. tatarinowii in 1966, holotype.

Anacardiaceae

Rhus trichocarpa Miq., Ann. Mus. Lugd.-Bat. 2: 84 (1865); Prolus. Fl. Jap.: 16 (1865).
Present treatment. Rhus trichocarpa Miq.: Hara, Enum. Sperm. Jap. 3: 66 (1954). Iwatsuki in Fl. Jap. 2c: 58 (1999).
Type material. Siebold s.n. [897. 363-1077] (L 0421974), holotype.

Aceraceae

Acer buergerianum Miq., Ann. Mus. Lugd.-Bat. 2: 88 (1865); Prolus. Fl. Jap.: 20 (1865).
Present treatment. Acer buergerianum Miq.: H. Hara, Enum. Sperm. Jap. 3: 98 (1954).
Type materials. In Iaponia, sponte crescens, Keiskeo (prob. in Owari) & Siebold s.n. [899. 296-144] (L 0329805), lectotype; [908. 268-1304] (L 0329803), syntype.
Other materials. Buerger [908. 269-594] (L 0104095); [899. 300-43, duplicate of 908. 269-594] (L 0104094).
Acer sieboldianum Miq., Ann. Mus. Lugd.-Bat. 2: 87 (1865); Prolus. Fl. Jap.: 19 (1865).
Present treatment. Acer sieboldianum Miq.: Hara, Enum. Sperm. Jap. 3: 115 (1954). Ogata in Fl. Jap. 2c: 65 (1999).
Type material. In montibus. Figo [Op een berg in Figo], Keiske s.n. [899. 300-34] (L 0170913), holotype.
Other material. Keiske s.n. [908. 268-1089] (L 0014697).
Negundo? nikoense Miq., Ann. Mus. Lugd.-Bat. 2: 90 (1865); Prolus. Fl. Jap.: 22 (1865).
Present treatment. Parthenocissus tricuspidata (Siebold & Zucc.) Planch.: Hara, Enum. Sperm. Jap. 3: 136 (1954).
Type material. In montibus Niko rarissimum; arbor valida [Keiske s.n.] [908. 268-1302] (L 0013700), with Hara's determination as Parthenocissus tricuspidata (Siebold & Zucc.) Planchon in Aug. 1954, holotype.

Sabiaceae

Meliosma oldhamii Miq., Ann. Mus. Lugd.-Bat. 3: 94 (July 2, 1867); Prolus. Fl. Jap.: 258 (1867).
Meliosma oldhamii Maxim.: Hara, Enum. Sperm. Jap. 3: 120 (1954).
Present treatment. Meliosma arnottiana (Wight) Walp. subsp. oldhamii (Maxim.) H. Ohba var. oldhamii: Ohba in Fl. Jap. 2c: 78 (1999).
Type material. Archipelagus Coreanus, Oldham no. 183 [908. 204-417] (L 0013914), holotype.

　This is a later homonym of Meliosma oldhamii Maxim. in Bull. Acad. Imp. Sci. Saint-Péacute; tersbourg 12: 64 (June 26, 1867) based on the same type.

Meliosma tenuiflora Miq., Ann. Mus. Lugd.-Bat. 3: 93 (1867); Prolus. Fl. Jap.: 257 (1867).
Present treatment. Meliosma tenuis Maxim.: Ohba in Fl. Jap. 2c: 78 (1999).
Type material. Keiske s.n. [908. 204-402] (L 0013842), holotype.

Balsaminaceae

Impatiens textorii Miq., Ann. Mus. Lugd.-Bat. 2: 76 (1865); Prolus. Fl. Jap.: 8 (1865).
Present treatment. Impatiens textorii Miq.: Hara, Enum. Sperm. Jap. 3: 123 (1954). Akiyama in Fl. Jap. 2c: 80 (1999).
Type materials. Textor s.n. [903. 280-104] (L 0420994), holotype; [903. 280-102] (L 0420996), isotype; [903. 280-103] (L 0420995), isotype.
Other materials. Siebold s.n. [903. 280-101] (L 0326359). Pierot s.n. [903. 280-106] (L 0326356).

Aquifoliaceae

Ilex* argutidens Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 84 (1866); Ann. Mus. Lugd.-Bat. 3: 104 (1867); Prolus. Fl. Jap.: 268 (1867).
Ilex serrata Thunb. var. argutidens (Miq.) Rehder in Bailey, Cycl. Amer. Hort. 2: 798 (1900). Koidz., Fl. Symb. Orient-Asiat.: 8 (1930). Hara, Enum. Sperm. Jap. 3: 78 (1954).
Present treatment. Ilex serrata Thunb. var. serrata: Ohba & Akiyama in Fl. Jap. 2c: 90 (1999).
Type material. Siebold s.n. [904. 134-314] (L 0015498), holotype.
Ilex* buergeri Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 84 (1866); Ann. Mus. Lugd.-Bat. 3: 106 (1867); Prolus. Fl. Jap.: 270 (1867).
Present treatment. Ilex buergeri Miq.: Hara, Enum. Sperm. Jap. 3: 68 (1954). Ohba & Akiyama in Fl. Jap. 2c: 88 (1999).
Type material. Buerger s.n. [904. 134-52] (L0015484), holotype.
Ilex macropoda Miq., Ann. Mus. Lugd.-Bat. 3: 105 (1867); Prolus. Fl. Jap.: 269 (1867).
Present treatment. Ilex macropoda Miq.: Hara, Enum. Sperm. Jap. 3: 73 (1954). Ohba & Akiyama in Fl. Jap. 2c: 91 (1999).
Type materials. Buerger [904. 134-228] (L 0017889), holotype; [904. 134-231] (L 0017890), isotype.
Ilex oldhamii Miq., Ann. Mus. Lugd.-Bat. 3: 105 (1867); Prolus. Fl. Jap.: 269 (1867).
Present treatment. Ilex chinensis Sims: Hara, Enum. Sperm. Jap. 3: 68 (1954). Ohba & Akiyama in Fl. Jap. 2c: 85 (1999).
Type materials. Prope Nangasaki, Oldham no. 149 [904. 134-252] (not included in the database), lectotype (here chosen); no. 150, syntype [not found]; no. 151 [904. 134-259] (not included in the database), syntype. Siebold s.n. [904. 134-264] (L 0328766), syntype; [904. 134-263] (L 0328765), syntype. Mohnike s.n. [904. 134-257] (L 041456), syntype; [904. 134-258] (L 0421455), syntype.
Ilex* pedunculosa Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 83 (1866); Ann. Mus. Lugd.-Bat. 3: 106 (1867); Prolus. Fl. Jap.: 270 (1867).
Present treatment. Ilex pedunculosa Miq.: Hara, Enum. Sperm. Jap. 3: 75 (1954). Ohba & Akiyama in Fl. Jap. 2c: 86 (1999).
Type materials. Buerger s.n. [904. 134-275] (L 0326675), lectotype (here chosen); [904. 134-269] (L 0326672), isolectotype; [904. 134-271] (L 0326674), syntype.
Ilex* sieboldii Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 84 (1866); Ann. Mus. Lugd.-Bat. 3: 104 (1867); Prolus. Fl. Jap.: 268 (1867).
Present treatment. Ilex serrata Thunb.: Koidz., Fl. Symb. Orient.-Asiat.: 8 (1930). Hara, Enum. Sperm. Jap. 3: 77 (1954). Ohba & Akiyama in Fl. Jap. 2c: 90 (1999).
Type materials. Buerger s.n. [904. 134-333] (L 0328834), lectotype (here chosen); [904. 134-327] (L 0329026), syntype. Siebold s.n. [904. 134-332] (L 0328835). Koidzumi wrote “this is that typical form of Ilex serrata” on the sheet.
Ilex* subpuberula Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 84 (1866); Ann. Mus. Lugd.-Bat. 3: 106 (1867); Prolus. Fl. Jap.: 270 (1867).
Present treatment. Ilex buergeri Miq.: Hara, Enum. Sperm. Jap. 3: 68 (1954). Ohba & Akiyama in Fl. Jap. 2c: 88 (1999).
Type material. Prope Nangasaki, Oldham no. 148 [904. 134-53] (L 0015483), holotype.
Ilex* subtilis Miq. in Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch. ser. 2, vol. 2: 84 (1866); Ann. Mus. Lugd-Bat. 3: 106 (1867); Prolus. Fl Jap.: 271 (1867).
Ilex serrata Thunb. var. subtilis (Miq.) Losen.: Koidz., Fl Symb. Orient.-Asiat.: 8 (1930).
Present treatment. Ilex serrata Thunb. (var. serrata): Hara, Enum. Sperm. Jap. 3: 77 (1954). Ohba & Akiyama in Fl. Jap. 2c: 90 (1999).
Type material. Siebold s.n. [904. 134-350] (L 0015499), holotype, I. serrata f. subtilis (Miq.) Ohwi det. Hara, Aug. 1954.

Celastraceae

Celastrus adenophylla Miq., Ann. Mus. Lugd.-Bat. 2: 85 (1865); 3: 104 (1867), in syn. of Ilex crenata; Prolus. Fl Jap.: 17 (1867).
Present treatment. Ilex crenata Thunb. (var. crenata): Hara, Enum. Sperm. Jap. 3: 69 (1954). Ohba & Akiyama in Fl. Jap. 2c: 83 (1999). [Aquifoliaceae]
Type material. Textor [904. 134-90] (L 0017885), holotype.
Celastrus ciliidens Miq., Ann. Mus. Lugd.-Bat. 2: 85 (1865); Prolus. Fl. Jap.: 17 (1865).
Present treatment. Celastrus flagellaris Rupr.: Hara, Enum. Sperm. Jap. 3: 79 (1954). Noshiro in Fl. Jap. 2c: 100 (1999).
Type material. Keiske no. 16 [899. 173-296] (L 0042887), holotype.
Euonymus japonicus Thunb. var. β radicans Miq., Ann. Mus. Lugd.-Bat. 2: 86 (1865); Prolus. Fl. Jap.: 18 (1865).
Present treatment. Euonymus fortunei (Turcz.) Hand.-Mazz. var. radicans (Miq.) Rehder: H. Hara, Enum. 3: 84 (1954). Noshiro in Fl. Jap. 2c: 96 (1999).
Type materials. Siebold [899. 174-213] (L 0328795), holotype; [899. 174-214] (L 0328800), isotype. This specimen has a label with the handwritten “Evonimus (?) radicans, S”.
Euonymus oxyphyllus Miq., Ann. Mus. Lugd.-Bat. 2: 86 (1865); Prolus. Fl. Jap.: 18 (1865).
Present treatment. Euonymus oxyphyllus Miq.: Koidz., Symb. Fl. Orient-Asiat.: 13 (1930). Hara, Enum. Sperm. Jap. 3: 89 (1954). Noshiro in Fl. Jap. 2c: 99 (1999).
Type materials. In insula Nippon, Siebold s.n. [899. 174-208] (L 0017896), lectotype (here chosen); [899. 174-203] (L 0017894), syntype; [899. 174-204] (L 0017897), syntype. In monte haud procul ab Aso [Op een berg niet ver van Aso], Keiske s.n. [899. 174-206] (L 0042889), syntype. In vallibus montis Homan Dake ins. Kiusiu[Kiu Siu], Pierot [no. 94] [899. 174-207] (L 0017895), syntype.
Other material. Buerger s.n. [899. 174-205] (L 0170532).

Rhamnaceae

Rhamnella japonica Miq., Ann. Mus. Lugd.-Bat. 3: 30 (1867); Prolus. Fl. Jap.: 218 (1867).
Present treatment. Rhamnella franguloides (Maxim.) Weberb.: Hara, Enum. Sperm. Jap. 3: 126 (1954). Noshiro in Fl. Jap. 2c: 118 (1999).
Type materials. Prope Nangasaki, Oldham no. 165 [908. 190-1321] (not included in the database), lectotype (here chosen). In ins. Kiusiu prov. Higo, Maximowicz s.n. [908. 190-1430] (not included in the database), syntype.
Rhamnus buergeri Miq., Ann. Mus. Lugd.-Bat 3: 31 (1867); Prolus. Fl. Jap.: 219 (1867).
Present treatment. Rhamnus japonica Maxim, var. decipiens Maxim.: Hara, Enum. Sperm. Jap. 3: 128 (1954).
Type materials. Buerger s.n. [908. 190-1413] (L 0104246), holotype; [908. 190-1372] (L 0104244), isotype; [908. 190-1400] (L 0104247), isotype.
Other materials. Buerger [908. 190-1397] (L 0104245); [908. 190-1440] (L 0104243).

　The specimens, [908. 190-1397] (L 0104245) and [908. 190-1440] (L 0104243), have matured fruits and seem to be collected in later season from the same tree or population from which the type was collected.

Vitaceae

Vitis** inconstans Miq., Ann. Mus. Lugd.-Bat. 1: 91 (1863); 2: 157 (1866); Prolus. Fl. Jap.: 89 (1866).
Present treatment. Parthenocissus tricuspidata (Siebold & Zucc.) Planch.: Hara, Enum. Sperm. Jap. 3: 136 (1954). Ohba in Fl. Jap. 2c: 123 (1999).
Type materials. Thunberg in 1777 [897.347-297] (L 0144331), lectotype (here chosen). In saxetis & ruderibus circa urbem Nangasaki insulae Kiusius, Siebold, syntype [not found].

　When Miquel published this species in Ann. Mus. Lugd.-Bat. 1: 91, 1863, he cited two specimens: “Iaponia, in muris Decimae & alibi, teste Thunberg” and “In saxetis & ruderibus circa urbem Nangasaki insulae Kiusius: Siebold.” So we consider that these two specimens are syntypes. Later he cited three specimens in Prolusio (in Ann. Mus. Lugd.-Bat. 2: 157, 1866): “Thunber a. 1777 tetexit”, “in monte Takatika in Hiugo invenit (“frutex repens”) Siebold” and “in saxetes & ruderibus circa Nangasaki Pierot.” We could not find the specimen collected by Siebold in Nangasaki.

Other materials. Siebold s.n. [897. 347-304] (L 0104271). Nakai determined this as mixture of Ampelopsis brevipedunculata and Quinaria thunbergii.; In monte Takatika[Takatiho] in Hiugo[Hiuga] (frutex repens), Siebold [Keiske? s.n.] [897. 347-281] (L 0170908), “Acer Negundo” det. Nakai. In montibus Witzo, (Udagawa) Jooan s.n. [897. 347-300] (L 0328840), as V. flexuosa by Nakai. In saxetis & ruderibus circa Nangasaki, Pierot s.n. [897. 347-299] (L 0104270).

Tiliaceae

Tilia cordata Mill. var. japonica Miq., Ann. Mus. Lugd.-Bat. 3: 18 (1867); Prolus. Fl. Jap.: 206 (1867).
Present treatment. Tilia japonica (Miq.) Simonk.: Hara, Enum. Sperm. Jap. 3: 142 (1954). Noshiro in Fl. Jap. 2c: 129 (1999).
Type materials. In declivibus[declitate] montis[montium] Sata Toge, prope oppidum Kifura Sawa ins. Nippon, Pierot [no. 568] [908. 254-81] (L 0175352), lectotype (here chosen). In ditione Figo, Keiske s.n. [908. 254-91] (L 0170926), syntype. In montibus Hakone, Sugerok [no. 89] [908. 254-94] (L 0170925), syntype.

Malvaceae

Hibiscus (Abelmoschus) japonicus Miq., Ann. Mus. Lugd.-Bat. 3: 19 (1867); Prolus. Fl. Jap.: 207 (1867).
Present treatment. Abelmoschus manihot (L.) Medik. subsp. manihot: Hara, Enum. Sperm. Jap. 3: 145 (1954). Ohba in Fl. Jap. 2c: 137 (1999).
Type material. Siebold s.n. [908. 135-244] (L 0012901), holotype, as “Abelmoschus manihot (L.) Medik. subsp. manihot” by Borssum Woalkes in 1989, on the sheet.
Malvastrum**** ruderale (B1.) Miq., Flora van Nederlandsch Indië 1(2): 138 (1858); Ann. Mus. Lugd.-Bat. 3: 19 (1867); Prolus. Fl. Jap.: 207 (1867).
Present treatment. Malvastrum coromandelianum (L.) Garcke: Ohba in Fl. Jap. 2c: 138 (1999).
Type materials. Siebold [908. 137-109] (L 0327571), lectotype; [908. 137-221] (L 0327572), syntype.

Thymelaeaceae

Daphne kiusiana Miq., Ann. Mus. Lugd.-Bat. 3: 134 (1867); Prolus. Fl. Jap.: 298 (1867).
Present treatment. Daphne kiusiana Miq.: Hara, Enum. Sperm. Jap. 3: 231 (1954). Murata in Fl. Jap. 2c: 147 (1999).
Type materials. In montibus ins. Kiusiu, Siebold s.n. [908. 239-1019] (L 0010158), holotype; [908. 239-1039] (L 001059), isotype.
Daphne odora Thunb. [var.] β marginata Miq., Ann. Mus. Lugd.-Bat. 3: 133 (1867); Prolus. Fl. Jap.: 297 (1867).
Daphne odora Thunb. f. marginata Makino in Bot. Mag. Tokyo 23: 69 (1909): Hara, Enum. Sperm. Jap. 3: 233 (1954).
Present treatment. Daphne odora Thunb.: Murata in Fl. Jap. 2c: 147 (1999).
Type materials. Siebold s.n. [908. 239-1053] (L 0010160), holotype; [908. 239-1063] (L 0010161), isotype.
Wikstromia canescens Meissn. var. gampi (Meissn.) Miq. forma japonica Miq., Ann. Mus. Lugd.-Bat. 3: 133 (1867); Prolus. Fl. Jap.: 297 (1867).
Present treatment. Diplomorpha trichotoma (Thunb.) Nakai.
Type materials. Siebold s.n. [944. 238-20] (L 0104379), lectotype, det. as W. japonica Miq. by Miquel; [944. 238-21] (L 0104372), syntype, as W. japonica Miq.; [908. 239-1462] (L 0104380), syntype; [908. 239-1463] (L 0104378), syntype. Keiske s.n. [908. 239-1450] (L 0420496), syntype. Buerger s.n. [908. 239-1466] (L 01404381), syntype. [In planitie] Prope pagum Meta Bara insulae Kiusiu, Pierot [no. 806] [908. 239-1464] (L 0104375), syntype; [908. 239-1465] (L 0104379), isosyntype. Prope Nangasaki, Mohnike s.n. [908. 239-1468] (L 0420495), syntype. Prope Jokohama, Maximowicz, syntype [not found].
Other material. Buerger [908. 239-1469] (L 0104377).

Violaceae

Viola keiskei Miq., Ann. Mus. Lugd.-Bat. 2: 153 (1866); Prolus. Fl. Jap.: 85 (1866).
Present treatment. Viola keiskei Miq.: Hara, Enum. Sperm. Jap. 3: 205 (1954). Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 170 (1999).
Type material. Keiske s.n. [908. 268-1001] (L 0175365), holotype.
Viola pinnata L. var. dissecta Turcz. forma a. trisectopinnatifida Miq., Ann. Mus. Lugd.-Bat. 2: 152 (1866); Prolus. Fl. Jap.: 84 (1866).
Present treatment. Viola chaerophylloides (Regel) W. Becker var. chaerophylloides: Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 175 (1999)..
Type materials. In locis arenosis prope prom. [In locis arenosis promontorii] Nomo Saki [in Kiu Siu], Pierot [no. 410] [908. 268-959] (L 0325791), lectotype (here chosen). Ad portum Hamilton, Oldham s.n. [908. 268-800] (not included in the database), syntype, as “var. trisectopinnata fide Miq.” Buerger s.n. [908. 268-780] (L 0325794), syntype; [908. 268-960] (L 0325792), syntype; [908. 268-979] (L 0325793), syntype.
Viola pinnata L. var. dissecta Turcz. forma b. triphylla Miq., Ann. Mus. Lugd.-Bat. 2: 152 (1866); Prolus. Fl. Jap.: 84 (1866).
Present treatment. Viola eizanensis (Makino) Makino: Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 175 (1999).
Type materials. Buerger [908. 268-790] (L 0325798), lectotype (here chosen); [908. 268-980] (L 0325795), syntype.
Other material. A Siebold collection by a Japanese [908. 268-958] (L 0325796).
Viola pinnata L. var. dissecta Turcz. forma c. pedata Miq., Ann. Mus. Lugd.-Bat. 2: 152 (1866); Prolus. Fl. Jap.: 84 (1866).
Present treatment. Viola chaerophylloides (Regel) W. Becker var. chaerophylloides: Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 175 (1999).
Type material. In valle Karnioan [Mamisaka] in Higo, in regione inter vulcanum Aso & Sukejama, Siebold s.n. [908. 268-799] (L 0325786), holotype.
Other materials. Siebold s.n. [908. 268-779] (L 0325787); [908. 268-788] (L 0325789).
Viola prionantha Bunge var. latifolia Miq., Ann. Mus. Lugd.-Bat. 2: 153 (1866); Prolus. Fl. Jap.: 85 (1866).
Present treatment. Viola phalacrocarpa Maxim.: Hara, Enum. Sperm Jap. 3: 213 (1954).
Type material. Siebold [not found].
Viola sylvestris Lam. forma b. deltoidea Miq., Ann. Mus. Lugd.-Bat. 2: 154 (1866); Prolus. Fl. Jap.: 86 (1866).
Present treatment. Viola grypoceras A. Gray. (syn. nov.)
Type materials. Siebold s.n. [908. 269-383] (L 0325802) (right and lower only), lectotype (here chosen). Buerger s.n. [908. 269-403] (L 0325803), syntype.

　The lectotype specimen, Siebold s.n. [908. 269-383], is V. grypoceras. The syntype specimen, Buerger s.n. [908. 269-403], consists of four plants: the lower and middle right are V. grypoceras, the upper and middle left are V. obtusa (Miq.) Makino.

Viola sylvestris Lam. forma c. exilis Miq., Ann. Mus. Lugd.-Bat. 2: 154 (1866); Prolus. Fl. Jap.: 86 (1866).
Present treatment. Viola grypoceras A. Gray var. exilis (Miq.) Nakai: Maekawa in Hara, Enum. Sperm. Jap. 3: 202 (1954), as “V. sylvestris Kitaib. var. c. exilis Miquel”. Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 180 (1999).
Type materials. Buerger s.n. [908. 269-384] (L 0424518), lectoytpe. Prope Nangasaki, Siebold s.n. [908. 269-363] (L 0424517), syntype.

　The lectotype specimen, Buerger s.n. [908. 269-384], consists of five plants. At least upper left and middle are V. obtusa; the upper right and lower right represent V. grypoceras var. exilis. The lower left seems to belong to the same variety.

Viola sylvestris Lam. forma d. obtusa Miq., Ann. Mus. Lugd.-Bat. 2: 154 (1866); Prolus. Fl. Jap.: 86 (1866).
Present treatment. Viola obtusa (Makino) Makino: Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 182 (1999).
Type material. Buerger s.n. [908. 269-415] (L 0325808), holotype. This consists of three plants and upper one is V. grypoceras.
Other materials. Siebold s.n. [908. 269-365] (L 0325809), this consists of 11 individuals of V. obtusa. Siebold s.n. [908. 269-395] (L 0325807), this consists of six individuals of V. grypoceras.
Viola sylvestris Lam. forma e. ovato-oblonga Miq., Ann. Mus. Lugd.-Bat. 2: 154 (1866); Prolus. Fl. Jap.: 86 (1866).
Present treatment. Viola ovato-oblonga (Miq.) Makino: Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 181 (1999).
Type materials. Siebold s.n. [908. 269-375] (L 0325810) (left two only), lectotype (here chosen); [908. 269-370] (L 0325812), syntype.

　Siebold [908. 269-375] consists of five plants; the lower two are V. grypoceras with narrowly ovate or lanceolate deeply incised stipules. Siebold [908. 269-370] consists of two individuals; the left one is V. ovato-oblonga but the right is V. grypoceras.

Viola sylvestris Lam. forma f. acaulis Miq., Ann. Mus. Lugd.-Bat. 2: 154 (1866); Prolus. Fl. Jap.: 86 (1866).
Present treatment. Viola grypoceras A. Gray (var. grypoceras): Hara, Enum. Sperm. Jap. 3: 200 (1954). Akiyama, Ohba & Tabuchi in Fl. Jap. 2c: 179 (1999).
Type materials. Siebold s.n. [908. 269-392] (L 0325799), lectotype (here chosen). Buerger s.n. [908. 269-404] (L 0325801), syntype. Textor s.n. [908. 269-373] (L 0328097), syntype, this is V. grypoceras.
Other material. In margine viarum juxte urbem Nagasaki in Kiu Siu, Pierot no. 412 [908. 269-412] (L 0325800).

　This is a seashore type of V. grypoceras which is found around Nagasaki.