Lespedeza bicolor Turcz.

In Bull. Soc. Nat. Mosc. 13: 69 (1840)-Ledeb., Fl. Ross. 1: 715 (1842)-Maxin., Prim. Fl. Amur., 86 et 470 (1859); in Acta Hort. Petrop. 2: 355 (1873), incl. α. typica (excl. quoad specim. cit. 'Colitur circa templa Nagasaki floribus candidis') et β. intermedia Maxim. (incl. quoad specim. cit. Tatarinow), excl, r. Sieboldii-Regel in Garten-ftora 60: 270, t. 299 (1860); in Mém. Acad. Imp. Sci. St.-Pétersb. 7 sér. 4: 48 (1861)-Miquel, Ann. Bot. Ludg.-Bat. 3: 47 (1867) et Prol. Fl. Jap., 235 (1867), excl. quoad specim. cit. Buerger, Siebold (pro parte) et Textor-Fr. Schmid tin Mém. Acad. Imp. Sci. St.-Pétersb. 7 sér. 12: 38 (1868)-Koch, Dendrol., 1: 73 (1869)-Debeaux in Act. Soc. Linn. Bordeaux 31: 140 (1876), pro parte (incl. β. intermedia Maxim., excl. r. Sieboldii (Miq.) Maxim. et syn. L. Sieboldii Miq.)-Franchet & Savatier, Enum. Pl. Jap. 1: 101 (1875)-Franchet in Nouv. Arch. Mus. Paris, sér. 2, 5: 247 (1883); Pl. David. 1: 95 (1884)-Forbes & Hemsley in Journ. Linn. Soc. Bot. 23: 178 (1887), pro parte-Koehne, Deutsch. Dendrol., 343 (1893), pro parte (cf. Nakai, 1927)-Diels in Engl., Bot. Jahrb. 29: 415 (1901)-Matsumura in Bot. Mag. Tokyo 14: 51 (1902), pro parte, incl. var. intermedia Maxim. f. acutifolia Matsum., f. grandi-folia Matsum. et f. parvifolia Matsum.; Ind. Pl. Jap. 2(2): 267 (1912), pro parte-Ko-marov, Fl. Manshuriae 2: 599 (1904)-Schneider, Ill. Handb. Laubholz. 2: 112, fig. 70f, 71a-d (1907)-Nakai in Journ. Coll. Sci. Univ. Tokyo 26 (Fl. Koreana I): 155 (1909), pro parte; in Journ. Coll. Sci. Univ. Tokyo 32 (Fl. Koreana II): 467 (1911); in Bot. Mag. Tokyo 37: 73 (1923); Lesp. Jap. Korea, 63 (1927)-Pampanini in Nuov. Giorn, Bot. Ital. n. ser. 17: 398 (1910)-Makino in linuma, Somoku-dzusetsu (ed. Makino) 3: 997, pl. 19 (1912), pro parte-Schindler in Engl., Bot. Jahrb. 49: 583 (1913); in Sargent, Pl. Wils. 2: 112, pro parte-Sean, Trees Shrubs Brit. Isl. 2: 16 (1914), excl. var. alba Bean-Bailey, Stand. Cyclop. Hort. 4: 1845, fig. 2134 (1916), excl. "A white-fld. variety"-Makino & Nemoto, Fl. Jap., 734 (1925)-Render in Journ. Arn. Arb. 7: 173 (1926)-Vasil'ev in Komarov ed., Fl. U.S.S.R. 13: 379, pl. 20-1 (1928)-Kitagawa, Lineam. Fl. Mansh., 288 (1939); Neo-Liueam. Fl. Mansh., 405 (1979)-Ohwi, Fl. Jap., 678 (1953), excl. f. alba (Bean) Ohwi et syn. L. melanantha Nakai; Fl. Jap. rev. ed., 790 (1965), excl, f. alba (Bean) Ohwi et syn. L. melanantha Nakai; Fl. Jap. Eng. ed., 559 (1965), excl. f. alba (Bean) Ohwi; Fl. Jap. new ed., 790 (1975), excl. f. alba (Bean) Ohwi et syn. L. melanantha Nakai-Fu & Wang in Liou et al., Ill. Fl. Lig. Pl. N.-E. China, 342 (1955)-Fu in Acad. Sin. Bot., Ill. Important Chin. Pl. Leguminosae, 519 (1955)-Kitamura & Murata, Col. Ill. Herb. Pl. Jap. 2: 100 (1961)-Kung in Chinese Journ. Bot. 1: 22 (1963)-Hatusima in Mem. Fac. Agr. Kagoshima Univ. 6: 14 (1967), excl. syn. L. melanantha Nakai, L. melanantha Nakai var. longifolia Uyeki, L. melanantha Nakai f. rosea Nakai, L. homoloba Nakai var. higoensis T. Shimizu-Acad. Sin. Bot., Iconogr. Cormophyt. Sin. 2: 458, fig. 2646 1972)-Lee, in Bull. Seoul Nat. Univ. For. No. 2, 5 (1965), excl. f. alba (Bean) Ohwi; Ill. Fl. Korea, 468 (1979)-Ohashi in Satake et al., Wild Fl. Jap. Herb. 2: 205 (1982), excl. var. higoensis (T. Shimizu) Murata.
[Plates 10 & 11; Fig. 36]

Var. bicolor

L. bicolor Turcz. f. microphylla Miq. in Ann. Bot, Lugd.-Bat. 3: 47 (1867); Prol. Fl. Jap., 235 (1867), pro maj. parte.

L. bicolor Turcz. α. typica Maxim. in Acta Hort. Petrop. 2: 356 (1873), pro major parte.

L. bicolor Turcz. β. intermedia Maxim. in Acta Hort. Petrop. 2: 356 (1873), pro parte, excl. quoad specim. "Tschonoski s.n." et "Oldham 335"-Matsumurain Bot. Mag. Tokyo 16: 69 (1902), pro parte; Ind. Pl. Jap. 2: 267 (1912), pro parte.

L. bicolor Turcz. var. intermedia Maxim. f. acutifolia Matsum. in Bot. Mag. Tokyo 16: 69 (1902)-Ohwi, Fl. Jap., 678 (1953); Fl. Jap. rev. ed., 790 (1965); Fl. Jap. Eng. ed., 559 (1965); Fl. Jap. new ed., 790 (1975), ut "L. bicolor Turcz. f. acutifolia Mat-sum."

L. bicolor Turcz. var. intermedia Maxim. f. grandifolia Matsum. in Bot. Mag. Tokyo 16: 69 (1902).

L. bicolor Turcz. var. intermedia Maxim. f. parvifolia Matsum. in Bot. Mag. Tokyo 16: 69 (1902).

L. japonica (non L. H. Bailey) Schindl. in Engl., Bot. Jahrb. 46: Beibl. 106, 54 (1912), nom. nud.

L. bicolor Turcz. var. japonica Nakai in Bot. Mag. Tokyo 37: 73 (1923).

L. bicolor Turcz. var. sericea Nakai, Lesp. Jap. Korea, 66 (1927).

L. setiloba Nakai, Lesp. Jap. Korea, 68 (1927).

L. spicata Nakai et F. Maekawa in Bot. Mag. Tokyo 48: 52 (1934).

L. spicata Nakai et F. Maekawa f. acutifolia Nakai et F. Maekawa in Bot. Mag. Tokyo 48: 53 (1934).

L. Tobae H. Koidzumi in Journ. Pl. Iwateken 2: 75 (1937), typum non vidi.

L. ionocalyx Nakai in Journ. Jap. Bot. 15: 532 (1939).

L. bicolor Turcz. f. tomentella Hatusima in Mem. Fac. Agr. Kagoshima Univ. 6: 14 (1967).

L. bicolor Turcz. f. sericea (Nakai) Hatusima in Mem. Fac. Agr. Kagoshima Univ. 6: 14 (1967), non Matsum. (1902)

L. bicolor Turcz. f. Nakaiana Murata in Acta Phytotax. Geobot. 34: 129 (1983), nom. nov.

Japanese name. Yamahagi (Matsumura, 1902).

A perennial plant, 1.5-2 m high. Stems ascending, 1-3 cm in diameter; most of the terrestrial part lives a few years; branched in upper parts (a few branches coming from basal part of the main stem). Branches terete, ascending, sometimes dependent later; when young slightly angular with densely ascending appressed-sericeous (very rarely spreading) hairs (hairs 0.3 mm long, whitish), later glabrescent; distal parts die in winter, proximal parts (and the main stem itself) with axillary and adventitious winter buds; winter buds at lower parts mostly dormant.

Leaves trifoliolate, petiolate, stipulate, spirally arranged. Stipules free, linear-triangular-linear, 2-7 mm long, brownish, persistent. Petioles 0.5-4 cm long, hairy like the young branch. Rachides 5-15 mm long, similar to the petiole. Terminal leaf lets (at middle parts of branches) petiolulate; petiolules 1-3 mm long, swollen; lamina 2-6 cm long, 1-3.5 cm wide, entire, (broadly) elliptic-(broadly) ovate or obo-vate, round or cuneate at the base, obtuse, refuse, or acute at the apex (the apex itself with or without a point), upper surface glabrescent (except along the midrib) or sericeous (hairs 0.3-0.5 mm long) at anthesis, lower surface appressed-sericeous (hairs 0.2-0.3(-0.5) mm long); lateral ones similar to terminal but somewhat smaller.

Inflorescence axillary racemous, solitary, 2-7(-10) cm long including the peduncle, somewhat loosely 4-12-flowered; peduncles 1-3 cm long, hairy like the young branch. Primary bracts ovate, about 1 mm long, pubescent, persistent; secondary bracts similar to primary ones.

Flowers 9.6-12.6 mm long at anthesis; pedicels 0.8-3 mm long, terete, curved, tomentose. Bracteoles at the base of the calyx broadly to narrowly ovate or obtuse, 0.6-1.2 mm long, pubescent outside, glabrous inside, persistent. Calyx 3.1-4.7 mm long, tubulate, four-lobed near the middle part or lesser than or more than the middle part, densely appressed pubescent; tube 1.5-2.4 mm long; lobes subequal in length or the lower one longest; lateral ones 1.2-2.3 mm long, 0.9-1.3 mm wide, ovate or triangular-lanceolate, obtuse or acute at the apex but not acuminate; upper one slightly two-cleft or not.

Standard longer than keel-petal, keel-petal longer than (rarely equal to or very rarely slightly shorter than) wings (S>K>W). Standard red-purple inside, paler outside, whitish near the base, obovate, 9.3-11.8 mm long, 5.4-7.2 mm wide, round or retuse at the apex with or without a point, attenuated at the base without distinct claw, inflexed-auriculate near the base, spreading (or reflexing) in anthesis from the part of 1/3 from the base. Wings deeper red-purple than standard, 7.3-9.5 mm long with distinct claw; lamina narrowly obovate to obovate, 4.4-6.8 mm long, 2.0-2.5 mm wide, auriculate at upper basal part; claw 3.2-3.9 mm long, whitish. Keel-petal 8.6-9.8 mm long with distinct claw; lamina obovate, 5.4-6.0 mm long, 2.3-2.9 mm wide, paler than wings, deepest in the apical part; claw 3.4-4.3 mm long, whitish. Stamens 10, nearly the same length, 8.4-9.6 mm long, diadelphous. Anthers elliptic, shallowly retuse at the apex, ca. 0.5 mm long; before anthesis yellow. Pistil 8.5-9.7 mm long; ovary elliptic, 1.5-2 mm long at anthesis, pubescent, subsessile; style 7-8 mm long, pubescent at the basal part, glabrescent at the apical part.

Fruits compressed broadly elliptic to round, 5-7 mm long, 4-6 mm wide, subses sile, slightly to densely pubescent or nearly glabrous. Seeds reniform, ca. 3 mm long, ca. 2 mm wide.

Distr. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Korea, N. China, and Amur.

Voucher and representative Specimens.

Japan (Siebold s.n., L 908.118-1080 (right)); (Buerger s.n., GH).

HOKKAIDO. SOUYA. Isl. Rebun, Lake Kushu-ko (Midorikawa 123, TI); Mt. Rishiri-Fuji, Oshidomari (Tuyama, 7 Sept. 1974, TI); Is. Rishiri, Mt. Rishiri, Foot Oshidomari-guchi (Furuse 21579, TNS); Wakkanai (Hiroe 7656, TI); Esaahi (Yamanoi, 8 Aug. 1971, TNS). ABASHIRI. Kitami, Rubeshibe (Wilson 7382, A); Nokkeushi (Uno 21551 & 21480, A); Mon-betsu-gun, Takinoue-machi (Okamoto, 20 July 1952, KYO); Tokoro-gun, Tokoro-cho (Oka-moto, 25 Aug. 1963, KYO); Kitami-shi, Hokkousha (Mizushima, 4 Oct. 1945, TI); Tsubetsu-cho, Goban, alt. 250m (Murata 21180, KYO); Abashiri-shi, Mokoto (Tanaka, 7 Sept. 1944, KYO); Shari-gun, Sharimachi, Iwaobetsu-onsen (Kobayashi, 19 July 1967, MAK). ABASHI-RI-NEMURO. Shiretoko, Mt. Rausu-take (Okamoto, 24 Aug. 1958, KYO). KUSHIRO. Kushi-ro-shi, Otanoshige (Naruhashi 1225, KYO); Tomachise (Hara, Kurosawa & Tateishi, 1934, TI); Lake Akan-ko (Hara c.l907.b, TI); Kawakami-gun, Shibetcha-cho, Shibetcha Experi mental Forest of Kyoto Univ. (Koyama 1733, TNS). TOKACHI. Rikubetsu (Okamoto, 4 Sept. 1955, KYO); Kamiashoro (Okamoto, 7 Aug. 1959, KYO); Hiroo-gun, Mt. Rakkodake-Satsu-rakko, alt. 200-400 m (Koyama & Fukuoka 3484, TNS). HIDAKA. Horoman (Hara, Kuro sawa & Tateishi, 1939b, 1940, TI); Shizunai-gun, Noya, Petegari-sanso, alt. 100-400 m (Koyama & Fukuoka 3106, TNS); Horoizumi, Syoya, Ruran (Hara, Kurosawa, & Tateishi, 1935, 1936, TI). KAMAGATA. Kamiotoineppu (Koidzumi, 8 Sept. 1930, TNS); Asahigawa (Asahikawa) (Koidzumi, Aug. 1916, TI; Aug. 1917, KYO); Kamikawa-gun, Kamuikotan (Uno s.n., A). SORACHI. Yubari-shi, Oyubari (Kuramoto 8108101-8108106, TI); Sorachi-gun, Toudai-enshuurin (Inokuma 2775, TOFO). ISHIKARI. Toyohirakawa (Matsumura, 1 Aug. 1899, TI-Syntype of L. bicolor Turcz. var. japonica Nakai); Sapporo (Yatabe, Aug. 1880, TI; Tanaka s.n., A; Miyabe, Aug. 1885, A); Mt. Moiwa-yama (Nakai, 25 Aug. 1920, TI); Teine (Takeda, 26 Sept. 1908, TNS); Zenibako (Matsuda, July 1887, KYO). SHIRIBESHI, Suttsu, Benkei-misaki (Hara & Kurosawa, 2 Sept. 1977, TI); Yoichi-machi (Okuda, 19 Aug. 1960, TNS); Otaru (Matsumura, 26 July 1878, TI-Syntype of L. bicolor Turcz. var. intermedia Maxim. f. acutifolia Matsum.). IBURI. Tomakomai-shi, Numanohata, Utonai-numa (Koyama 5644, KYO); Mukawa-machi, near Shiomi Station (Ohba & Akiyama 3501-3510, TI); Muroran (ut "Mororan") (Sargent, Sept. 1892, A); Lake Touya-ko (Nakano, Oct. 1899, TNS). HIYAMA. Ichinowatari (Miyabe & Tokubuchi, 15 July 1890, TI-Lectotype of L. bicolor Turcz. var. intermedia Maxim. f. acutifolia Matsum. and lectotype of L. bicolor Turcz. var. japonica Nakai; KYO). OSHIMA. Hakodate (Maximowicz in 1861, A; L 908.118-1089); Ha-kodate-shi, Hakodate-yama (Midorikawa 1890, TI); Ohnuma-koen (Wilson s.n., A).

HONSHU. AOMOPI. Aomori (Kinashi, Oct. 1900, KYO); Hirosaki (Kinashi, Aug. 1904, KYO); Kamikita-gun, Shiwa-mura (Mizushima 31, TI); Higashitsugaru-gun, Takanozaki (Ku rosawa, 28 July 1959, TI); Masukawa-eirinsho-bunai (Kusaka 1854, TOFO); Tappizaki (Hara, 27 July 1959, TI); Nishitsugaru-gun, Tateoka-machi, Hirataki-numa (Furuse, 19 Sept. 1955, KYO), Nishitsugaru-gun, Takayama-inari, seaside (Hara, Kurosawa, & Tateishi, 15 Sept. 1974, TI); Shimokita Penin. Ushitaki-Nodai (Naito, 12 Aug. 1963, TNS); Shimokita Penin. Shiriya-Kukidounosaki (Ohashi 4308, 4579, TI); Shimokita-gun, Sai-mura, Harada (Mori, 22 Aug. 1957, MAK); Mt. Iwaki (Yatabe, 24 July 1880, TI-Syntype of L. bicolor Turcz. var. intermedia Maxim. f. grandifolia Matsum.); Mutsu-shi (Midorikawa 1950-1953, TI); Shimo-kita-gun, Wakinosawa-mura (Midorikawa 1897, 1898, 1899, TI); Shimokita-gun, Higashi-toori-mura, to Imooto-mura from Horobe (Furuse, 21 Sept. 1957, A). IWATE. Morioka (Toba 17, 647, TI; T. Saito, 2 Aug. 1932, KYO); Morioka-shi, Asagishi (Kikuchi, 24 Sept. 1967, TNS); Yuda (Toba 12, TI-Holotype of L. spicata Nakai et F. Maekawa, 18, TI-Holo-type of L. spicata Nakai et F. Maekawa f. acutifolia Nakai et F. Maekawa, 11, 19, TI; 10 Sept. 1932, KYO); Mt. Kunimi (Iwabuchi, 28 July 1941, TNS); Araya Sinmachi (Sugawara, 1 Aug. 1963, TUSG); Hienuki-gun, Uchikawame-mura, Dake (Kanai, 1 Aug. 1959, TI); Kuji-shi, Ookawame (Murata & Tabata, 28 July 1967, KYO); Kitakami-shi, Kurosawajiri-machi (Oi-kawa, 7 Aug. 1965, MAK); Otobe-mura (Kitamura & Murata 526, KYO); Izawa-gun, Shito-mae (Sugawara, 20 Aug. 1970, TUSG); Mt. Yakeishi-dake (Karizumi, 27 Aug. 1951, TNS); Kamihei-gun, Tsuchibuchi-mura (Hosoi 10988, TI); Iwate-gun, Tamayama-mura, Mt. Hi-megami-dake (Ogata 4145, TOFO); Shimohei-gun, Iwaizumi-cho, Ryuusendou (Shimizu, 14 Oct. 1957, KYO); Higashiiwai-gun, Mt. Tabashine-yama (Kikuchi, 17 Sept. 1967, TNS); Mt. Himekami (Muroi 4087, A); Kuzakai (Muroi 3971, A); Morioka, Mt. Iwayama (Muroi 4238, A). MIYAGI. Sendai (Iishiba, 23 Aug. 1928, TNS); Sendai-shi, Mt. Dainenji-yama (Suzuki, 10 Sept. 1952, TNS); Tamatsukuri-gun, Narugo-machi, Onikoube (Sugawara, Ohba et al. 708207, TI; TUSG); Katta-gun, Oogawara-machi, Mt. Zaou-san (Suzuki, 10 Sept., TNS); Shi-bata-gun, Kawasaki-machi, Kamafusa Dam (Ohba & Aldyama 573, TI); Shibata-gun, Kawa-saki-machi, Maekawa, alt. 200-250 m (Ohba & Akiyama 582-586, TI); Katsuta-gun, Zaou-machi, Tougatta (Ohba & Aldyama 593, TI); Ishinomaki-shi, Tashiro-jima (Suzuki, 26 Aug. 1956, TNS); Natori-gun, near Okunikkawa Station (Kanai, 7 Sept. 1959, TI); Kami-gun, Ka-miishi-mura (Suzuki, 18 Sept. 1936, TI); Kami-gun, Nakaniida-machi, Naruse River (Naito & Ishikawa, 1 Sept. 1978, TUSG). AKITA. Mt. Tegata-yama (Muramatsu, 30 Sept. 1930, TI); Kitaakita-gun, Yamase-mura (Furuie, 26 Aug. 1951, TNS); Oomagari-shi, Yotsuya-mura (Kobayashi, 29 Aug. 1936, TNS); Kawabe-gun, Yuuwa-mura, Tsubakigawa (Henmi, 31 July 1965, TNS); Yokote-shi, Mt. Mitake-san (Nakamura, Aug. 1959, MAK); Hiraka-gun, Masuda-machi (Ishida, 22 Aug. 1959, MAK); Kitaakita-gun, Nakamura-Mt. Moriyoshi-yama (Ohasbi 4787, TI); Oga-shi, Kitaura, Nishimizuguchi (Fujii 560089, TI); Noshiro-shi, Oto-mo-numa (Henmi, 23 Aug. 1963, TNS); Yuri-gun, Kisakata-machi, Mt. Chokai-san, en route from Kisakata to Hokodate, alt. ca. 460 m, 520 m, 560 m, & 610 m (Kaji, 31 July 1982, TI); Oga, Mayama-Kamimayama (Inokuma 3107, TOFO). YAMAGATA. Yamagata-shi (Yuuki 3204, TI); Tobishima (Mori, 8 July 1962, MAK) ; Yunohama (Yuuki 3348, TI); Atsumi (Yuuki, 30 July 1936, TNS); Yonezawa-shi, Bansei (Katou, 18 Aug. 1935, KYO); Nishiokitama-gun, Oguni-machi, Taruguchi-Takikura (Ohba & Akiyama 2699-2703, TI); Nishiokitama-gun, Oguni-machi, Ichinosawa-Ashimizunakasato (Ohba & Akiyama 2686-2688, TI); Kitamura-yama-gun, Tateoka-machi, Komatsu kannon-Higashisawa (Kanai, 8 Sept. 1959, TI); Cho-kaizan (ut "Chokaitan") (Okubo, 29 July 1887, TI-Lectotype of L. bicolor Turcz. var. intermedia Maxim. f. grandifolia Matsum. and syntype of L. bicolor Turcz. var. japonica Nakai); Nishitagawa-gun, Mt. Maya-zan (Mori, 18 Aug. 1959, MAK); Yonezawa-shi, Onogawa, near Sasahara, alt. ca. 400 m (Ohba & Akiyama 2708-2711, 2712-2714, TI); Odate, Hadachi, Motaisan (Sugawara, Saito, & Sato, 3 Sept. 1969, TUSG); Ginzan-onsen (Okuyama 24143, TNS); Karuizawagoe (Okuyama 24142, TNS); Minamiokitama-gun, Mt. Azuma, Niitakayu (Uno, 4 Aug. 1952, A); Mt. Atsumi (Ikegami 17378, A). FUKUSHIMA. Yomogita-mura (Imai, 28 Aug. 1934, TNS); Mt. Bandai-san (Tashiro, 7 Aug. 1937, KYO); Kitaaizu-gun, Higashi-yama, Mt. Senakaaburi-yama (Hara & Kurosawa, 31 July 1957, TI); Aizuwakamatsu, Sea-buriyama, alt. 700 m (Yamazaki 6604, TI); Yama-gun, Inawashiro-machi, near Kawakami, alt. 700-750 m (Ohba & Akiyama 2724-2745, TI); Aidzu (Yatabe & Matsumura, Aug. 1879, TI-Syntype of L. bicolor Turcz. var. intermedia Maxim. f. acutifolia Matsum.); Aidzu, Yu-moto (Yatabe & Matsumura, 4 Aug. 1879, TI-Syntype of L. bicolor Turcz. var. intermedia Maxim. f. grandifolia Matsum.); Kawanuma-gun, Aidzubange-machi, Toudera-Ketanomiya (Ohba & Akiyama 1820-1828, TI); Date-gun, Yanagawa-machi, Mt. Togarimori-yama (Iga-rashi, 9 Aug. 1961, TNS); Fukushima-shi, Tsuchiyuonsen-cho, Tsuchiyu-Onuma, alt. 500-650 m (Kurosaki 13402, SHO); Mt. Shirabu-san (Okamoto, 4 Sept. 1938, KYO); Akaiya-chi, alt. 500m (Yamazaki, 11 Aug. 1962, TI); Adachi-gun, Ohtama-mura, Adatara-onsen (Kanai & Tateishi 1141, TUSG); Kooriyama-shi, Nakada-machi, Shitaeda, alt. ca. 400 m (Ohba & Akiyama 2799-2801, TI); Kooriyama-shi, Nakada-machi, Shitaeda, alt. ca. 400 m (Ohba & Akiyama 2802-2804, 2805, TI); Minamiaidzu-gun, Shimogo-machi, Ono, alt. 400-500 m (Ohba & Akiyama 1929-1935, TI); Kooriyama-shi, near Nishita-machi, Sanchome, alt. 200-300 m (Ohba & Akiyama 2761-2773, TI). GUNMA. Minakami (Hisauchi, 5 Aug. 1933, TI; 20 Aug. 1933, TNS); Tanigawa-onsen (Maekawa 58-A34, TI); Mt. Shirane-san (Sakurai, Sept. 1885, TNS); Mt. Myogi (Ishii, 13 Sept. 1967, TNS); Mt. Haruna-san (Ohwi, 6 Sept. 1950, A; TI; TNS); Mt. Arafune-yama-Hitsuse (Kimura, 6 Oct. 1953, TI); Kouzu-bokujo (Okuyama, 22 July 1940, TNS); Ushi-gun, Iriyama-touge (unknown collector, 13 Sept. 1934, TI); Azuma-gun, Shinkazawa (Kanai, 19 July 1958, TI); Shimizu-touge (Sato, 23 July 1933, TNS); Tone-gun, Kawaba-mura, Mt. Joshu-Hotaka-yama, Sainokawara (Kanai, 5 July 1957, Tl); Tone-gun, Katashina-mura, Tokura (Makino in 1929, KYO); Tone-gun, Katashina-mura, nr. Lake Maru-numa, alt. ca. 1500 m (Ohba & Akiyama 1496-1507, TI); Tone-gun, Tone-mura (Ohba & Akiyama 1516-1519, TI); Tone-gun, Shirasawa-mura, Namae alt. ca. 550 m (Ohba & Akiyama 1520-1523, TI); Tone-gun, Minakami-machi, Yubiso, alt. ca. 600 m (Ohba & Akiyama 1575-1576, TI); Matsuida (Wakana, 26 Sept. 1954, TNS); Maebashi-shi, Taguchi, Mt. Tachibana-yama (Shiobara, Aug. 1928, TNS); Tano-gun, Ueno-mura, Fujidoo, alt. 750-900 m (Murata, Ohba, & Midorikawa 6262, Tl); Kamizawa, alt. 1000 m (Wilson in 1914, A). TOCHIGI. Shiobara (Momiyama 1099, 1200, 1201, 1202, TI); Oku-shiobara (Suzuki, 9 Aug. 1951, A); Shioya-gun, Kuriyama-mura, Yunishigawa (Kubota, 31 Aug. 1965, TNS); Nikko (Sargent, 2 Sept. 1892, A); Nikko-shi, Arasawa-machi, alt. 650-700 m (Ohba & Aki yama 606 & 608, TI). IBARAKI. Tsukuba (Owatari, 21 July 1895, TI-Syntype of L. bicolor Turcz. var. intermedia Maxim. f. grandifolia Matsum. and syntype of L. bicolor Turcz. var. japonica Nakai); Shida-gun. (Hitachi) (Matsumura s.n.TI-Holotype of L. bicolor var. intermedia f. parvifolia Matsum.); Ishioka-shi, Ishioka-machi (Tsurumachi, 11 Sept. 1911, MAK). SAITAMA. Hannou (Hisauchi, 24 Sept. 1919, TI); Mt. Nobi-yama (Miyazaki in 1933, TNS); Niiza-machi, Katayama (Ohashi 9094, 9095, A; TI); Mt. Arashiyama (Hisauchi, 17 Sept. 1933, TI; TNS). CHIBA. Kashiwa (Okuyama 21279A, B, TNS); Kounodai (Makino, Sept. 1894, MAK); Inba-gun, Shimoshidzu-mura (Makino, 10 Sept. 1895, MAK); Teganuma (Makino in 1924, MAK); Inbanuma, Usui (Hisauchi, 23 Sept. 1933, TI); Ooami (Yamazaki, 6 Sept. 1956, TI); Shisui (Okuyama 9945, TNS); Ichikawa-shi, Ohmachi (Ohba 626905, TI). TOKYO. Inokashira (Nakai in 1928, TI); Ooi-futou (Midorikawa 1295, TI); Shibuya-ku, Yoyoki (Makino, Sept. 1900, KYO); Setagaya-ku, Chitose (Makino, 13 Oct. 1935, KYO); Suginami-ku, Iogi (Makino, 15 Sept. 1932, KYO); near Sekido (Mizushima, 15 Sept. 1946, TI); Chofu-near Ikou-ji (Mizushima 2458, TI); Kitatama-gun, Shinko-ganei (Kanai, 10 July 1949, TI); Hino-shi, Minamitaira (Ohba 1329, 1347, TI); Hikida-Ohme (Mizushima 422, A; TI); Nishitama-gun, Itsukaichi-machi, Yokosawa (Ohba 1785, 1832, TUSG). KANAGAWA. Mukogaoka, alt. 50 m (Suzuki 331, A); Mizonokuchi (Hisauchi, 6 Sept. 1931, TI); Noborito (Hisauchi, 12 Sept. 1932, TI); Kawasaki-shi, Ikuta (Makino, 16 Sept. 1948, KYO); Fujisawa-shi, Kameino, Mutsuai, Exp. Forests, Nippon Univ. (Takada, 6 July 1964, TNS); Ashigarashimo-gun, Hakone-machi, Sengokuhara (Ohba & Akiyama 648, TI); Hakone, Miyagino-mura (Mizushima 17, A). NIIGATA. Mt. Hachikoku-san (Ikegami, 23 Sept. 1944, TNS); Mt. Gozu-yama (Noda, 27 Aug. 1957, TNS); Ojiya, alt. 150 m (Yamazaki 8989, A; TI); Higashikanbara-gun, Tsugawa, Suwatoge, alt. 400 m (Yamazaki 255, TI); Kitakanbara-gun, Nakaura-mura (Kitakanbara-gun, Toyoura-mura) (Goto s.n., MAK); Mi-namikanbara-gun, Shitada-mura, Taya (Ohba & Akiyama 1747-1756, TI); Nakakanbara-gun, Muramatsu-machi, Maki (Ohba & Akiyama 1759, 1762, TI); Kunigami-mura, Mt. Kuni-gami-yama (Eguchi, Sept. 1951, TI); Nishikambara-gun, Mineoka (Ikegami 16443, A); Nii-gata-shi, Yamanoshita (Ikegami 9414, TNS); Nagaoka-shi, (Kusumi, 23 Sept. 1972, TNS); Kita-uonuma-gun, Kawaguchi-machi, alt. ca. 100 m (Ohba & Akiyama 1586-1593, 1594-1612, TI); Koshi-gun, Yamakoshi-mura (Ohba & Akiyama 1666-1677, 1690, TI); Joetsu-shi, Kasu-gayama-joshi, alt. 50 m (Ohba & Akiyama 467-468, TI); Kamo-shi, Nishiyama (Ohba & Akiyama 1757-1758, TI); Shibata-shi, w. foot of Mt. Iide, Higashiakatani-Yunohira Spa (Fukuoka & Konta, 18 July 1964, KYO); Minamiuonuma-gun, Mikuni-mura, Hiuchi Pass, alt. 920 m (Yamazaki, Kanai, Hasegawa, & Ohkubo 9036, A; TI); Minamiuonuma-gun, Yu-zawa-machi, near Asagai, alt. ca. 900 m (Ohba & Akiyama 1578-1585, TI); Nishikubiki-gun, Aomi-cho, Mt, Kurohimc (Hiroe 14301, KYO); Isl. Sado, Aikawa (Ikegami, 24 Sept. 1939, TNS). NAGANO. Akashina, alt. 600-1000 m (Wilson 7470, A); Mt. Hachibuse-yama (Momo-se, 24 Aug. 1933, TI); Mt. Kiso-koma, Naidaijo, alt. ca. 1400 m (Okamoto, 21 July 1950, KYO); Mt. Norikura, alt. 1850 m (Furuya, 15 Aug. 1950, A; TI); Mt. Nyugasa-yama, alt. 1000 m (Yamazaki, 25 Sept. 1954, TI); liyama-shi, Sekiya, alt. ca. 500 m (Ohba & Akiyama 493-495, TI); Nagano-shi, Susobana-KYO, alt. ca. 500 m (Ohba & Akiyama 498, 499, 500, 503-505, 506, TI); Kamiminochi-gun, Togakushi-mura, Toyooka, alt, 600-800 m (Ohba & Akiyama 513, 515, TI); Kamiminochi-gun, Kinasa-mura, Kinasa, alt. 700-800 m (Ohba & Akiyama 519, 522, 524, TI); Kamiminochi-gun, Ogawa-mura, nr. Meotoiwa, alt. ca. 1000 m (Ohba & Akiyama 526-528, TI); Kamiminochi-gun, Shinano-machi, near Lake Nojiriko, alt. ca. 700 m (Midorikawa 1005, TI); Oomachi-shi, Taira, nr. Lake Aoki, alt. ca. 750 m (Ohba & Akiyama 453-456, TI); liyama-shi, Sekiya, alt. ca. 500 m (Ohba & Akiyama 493-495, TI); Kitaazumi-gun, Hakuba-mura, Shirasawa, alt. ca. 900-950 m (Ohba & Akiyama 529-530, TI); Kitaazumi-gun, Hakuba-mura, Minekata, alt. 800-900 m (Ohba & Akiyama 531, TI); Minamiazumi-gun, Ariake (Hurusawa, Sept. 1939, TI); Sarashina-gun, Oooka-mura, Hika-geshiden (Mineura, 1 Aug. 1958, MAK); Komoro-shi (Makino in 1940, KYO); Nishichi-kuma-gun, Kiso-mura, Hosojima, alt. 1100 m (Kanai & Yamashita, 27 July 1964, TI); Wada-touge (Okamoto, 4 Aug. 1961, TNS); 2 km west of Kiyosato (Tuyama, 26 Sept. 1974, TI); Sugadaira, alt. ca. 1350 m (Teramoto, 27 Aug. 1947, TI); Chiisagata-gun, Shioda-cho, Bes-sho-onsen (Kitagawa 5362, KYO); Kamizawa (Okuyama, 17 April 1936, TNS); Mt. Ichinoji yama (Shiroishi, 25 Aug. 1935, TNS); Kutsukake-Kose, alt. 1000 m (Mizushima 10154, 10156, A; TI); Karuizawa, Mt. Yagasaki-yama (Kanai, 30 Aug. 1955, TI); Mt. Asama-yama (Ito, Aug. 1933, TI); Minamisaku-gun, Azusayama-Mikuniyama, alt. 1400 m (Murata, 7 Aug. 1958, KYO); Minamisaku-gun, Kitaaiki-mura, Shiroiwa, alt. 1000 m (Murata, Ohba, & Midorikawa 1216, TI); at the foot of Mt. Senjo-dake, Todai, alt. 1400 m (Uematsu, 24 Sept. 1949, TI); Ochikura (Hisauchi 1782, TI); Shimoina-gun, Iida, Mt. Hontakamori, alt. 800 m (Yamazaki & Asano 7524, TI); Shimoina-gun, Yamato-mura (Asano, Sept. 1963, TI); Shimoina-gun, Ooshika-mura, Nakao (Asano 9217, 4392, TI). YAMANASHI. Karisaka-touge, alt. 1500 m (Uematsu, 28 Sept. 1949, TI); Mt. Daibosatsu-toge (Okuyama 14618, TNS); Ki-takoma-gun, Masutomi-mura, Mt. Kinpu-san, alt. 1600 m (Kanai, 25 Sept. 1954, TI); Mita-ke, Shosen-KYO (Mizushima 189, TI); HIGAshiyamanashi-gun, Mt. Kentoku-san, alt. 1100 m (Fujita, 10 Oct. 1954, TI); Mt. Minobu-san (Oiwake-Akazawa) (Moriya, 9 Aug. 1966, TNS); Mt. Yatsu, Oizumi (Okuyama 12840, 12545b, TNS); N. side of Mt. Fuji, Yoshida-guchi, Umagaeshi-Ohishi-chaya, alt. 1400 m (Tateishi 374, TI); Fujiyoshida-shi, NE slope of Mt. Fuji, Nakanochaya Lodge, along Yoshida-guchi mountaineering path, alt. 1100 m (Konta, 24 Aug. 1976, MAK); W. side of Mt. Fuji (Ohba 69875, TI); Minamitsum-gun, Yamanaka Lake (Makino in 1936, KYO); Lake Kawaguchi-ko (Hisauchi 855, TI); Mt. Misaka (Kanai, 18 Sept. 1955, TI); at the foot of Mt. Houou-san (Hisauchi, 10 July 1938, TI); Mt. Houou-san, Aoki-kousen, alt. 1200 m (Yamazaki, 7 Sept. 1954, TI). SHIZUOKA. Inasa-gun, Uri-touge (Momiyama, 19 July 1951, TI); Sakuma-machi, Ohashi-bokujo (Okuyama 23266A, B, TNS). GIFU. Mt. Ibuki-yama, Sangou-me (Yamazaki, 3 Oct. 1959, TI); Ohgaki-shi, Naga-matsu (Yamazaki, 6 Aug. 1969, TI; TUSG); Ohno-gun, Asahi-mura, near Akigami Damsite (Onogi, 5 Sept. 1961, TNS); Mt. Norikura-dake, at the foot (Ito, 18 Aug. 1891, TNS); Toki-shi, Tajimi-machi (Shiota 7033, A); Toki-shi, Shimoishi-machi (Shiota 5393, A). AICHI. Tsukude-mura, Tashiro (Torii 6204, KYO; 6205, TUSG); Kamitsugu-mura, Orimoto-touge (Torii 6188, TUSG); Nishiura-machi (Torii 6179, TUSG); Hourai-machi, Ikeba (Torii, 29 July 1962, KYO); Minamishitara-gun, Mt. Tonbaku (Torii, 3 Nov. 1950, A); Kitashitara-gun, Inabu-machi, Kurakaitsu (Torii, 15 Sept. 1957, KYO); Toyone-mura, Itauba (Torii, 22 Sept. 1958, KYO); Kitashitara-gun, Miwa-mura, Uren-gawa (Yamazaki, 30 July 1957, TI); Shin-shiro-shi, Uri-touge (Torii, 4 Aug. 1963, TI); Toyohashi-shi (Torii 6185, TUSG); Seto-machi (Shiota, 21 Sept. 1930, KYO). TOYAMA. Shimoniikawa-gun, Unazuki-machi, Kurobe (Makino in 1935, KYO; MAK); Sakai-gawa (Hayashi, middle of Aug. 1930, KYO); Kaminiikawa-gun, Sasadzu (Shinno, 21 Sept. 1935, TNS); Nakaniikawa-gun, Kamiichi-machi, Asou (Ku-rosaki, Oct. 1966, KYO); Nei-gun, Kureha-mura, Itakura, HIME-jinja (Kotou, 29 Sept. 1940, TNS); TOYAMA-shi, west side of Mt. Kureha-yama (Shinno, 3 Oct. 1933, MAK). ISHIKAWA. Konosu (Kitamura, 22 July 1929, KYO); Ogi-machi (? Kogi-machi), Ohsawa (Hori 564, TOFO-Holotype of L. bicolor Turcz, f. tomentella Hatus. (ut "Kogi-machi")); Sosogi (Oku-yama 14697, TNS); Kashima-gun, Midukami-Sekidosan (Masamune 12315, MAK); Suzu-gun, Mt. Yamabushi-yama (Satomi, 8 Sept. 1960, KYO); Ishikawa-gun, Tsurugi-cho, west of Kuragadake (Fukuoka, 19 Oct. 1958, KYO); Hakui-gun, Mt. Houdatsu-san (Satomi, 29 Aug. 1961, KYO); Mt. Ioo, Miage-Nishiodaira (Fukuoka 4420, KYO; TNS); Mt. Kuraga-dake (Satomi, 13 Oct. 1953, TNS); Mt. Kigo (Satomi, 2 Sept. 1951, TI); Kanazawa-shi, Yuuhi-dera, alt. 50 m (Ohashi 9078, A; 9079, TI; 9088, A; TI). FUKUI. Ohno-shi, Rokuroshi (Murata & Shimizu, 22 Aug. 1954, KYO); Fukui-shi, Daian-ji (Ohashi, 14 Oct. 1964, TI); Mikata-gun, Mikata-cho, Tsunegami-misaki (Nakai, 26 July 1969, KYO); Yoshida-gun, Shi-moshigi-mura (Hori, 18 Sept. 1952, TNS); Itai-gun, Oshima-mura (Tashiro, 19 Sept. 1933, KYO; TNS); Imatate-gun, Mt. Hino-san (Hori, Aug. 1952, TNS). SHIGA. Mt. Ibuki-yama (Tashiro, 2 Sept. 1928, KYO); Ohtsu-shi, Mt. Ohsaka-yama (Hashimoto 11346, TNS). KYOTO. Mt. Kurama-yama (Hiroe 6445, TNS); Hukutiyama, Mutobe (Togasi, 21 Sept. 1949, TI); Mt. Kimio-yama (Araki 15492, TI); Mt. Ooeyama, at the summit, alt. 830 m (Murata, 11 Oct. 1967, KYO); Maidzuru-shi, Mt. Akaiwa-yama, Shimomidani-the summit, alt. 200-600 m (Murata 22379, KYO; TNS; TUSG); Yosa-gun, Futyu-mura (Murata 4950, KYO); Takeno-gun, Yasaka-cho, Noma, near Sukawa, alt. 200-300 m (Murata, 29 Sept. 1964, KYO). HYOGO. Mt. Rokkou-san, near Okuike (Yoshina, 5 Aug. 1934, KYO); Mikata-gun, Mt. Hachibuse-yama (MAKino, 9 Aug. 1937, KYO; Araki 14461, KYO; Nishihara, 19 Aug. 1937, TNS; Muroi 5432, A); Mikata-gun, Yohka (Muroi 5552, A); Yabe-gun, Sekinomiya-cho, Mt. Hachibuse, Oku-bo-Takamaru, alt. 700 m (Murata 21139, KYO; TI; TNS); Mikata-gun, Onsen-cho, Umigami-Mt. Ogino-seii (Murata 20749, KYO); Kinosaki-gun, Kasumi-cho, Yoroi, at the seaside (Murata, 23 Oct. 1966, KYO); Taki-gun, Hioki-mura, Kitajima-Kumobe-mura, Izumi (Araki 14561, KYO; TI); Nishihama-mura, Moroyose-kaigan (Hiratsuka 24, TI); Mt. Myoken-san (Tashiro, 26 Aug. 1932, TNS); Hikami-gun, Kasuga-machi, Kurotsuhou (Hosomi, 19 July 1936, KYO); Mt. Hyono-sen, at the foot (Makino, 8 Aug. 1937, KYO); Siso-gun, tikusa-mura, Nabegatani (Tagawa, 21 Aug. 1954, KYO); Siso-gun, Haga-cho, Tokura, alt. 500-700 m (Murata, 17 Aug. 1968, KYO; Murata & Hotta 13, TI; TNS); Sayou-gun, Nanko-cho, Mt. Hunakoshi-yama, alt. 300-650 m (Murata 33767, KYO); Kanzaki-gun, Ohkawauchi-cho, To-nomine Hills (Mizushima & KOYAMA, 17 Oct. 1966, TNS); Asako-gun (Araki 13737, KYO); Isl. Awaji, Mihara-gun, along the road extending north of Fukiage-cho (Fukuoka 9873, KYO). OKAYAMA. Niimi (Makino in 1935, MAK); Jobou-gun, Kayou-machi, Suzuyama (Nan-ba 9417, KAG); Tsuyama-shi, KOUCHIda (Nanba 9425, KAG); Katsuta-gun, Kitayoshino-mura (Hiratsuka 17, TI); Maniwa-gun, Yatsuka-mura (Tashiro s.n., KYO); Maniwa-gun, Yubara-cho, near Yubarako (Murata 27145 & 28 July 1962, KYO); Maniwa-gun, Mt. Kamihiruzen, Nakahukuda-the summit, alt. 700 m (Murata, 29 July 1962, KYO); Atsesu-gun, Ohnobe (Osaka, 15 Aug. 1929, KYO); Kawakami-gun, Hirakawa-mura (Tashiro, 13 Aug. 1932, KYO); Tsukubo-gun, Asabara, Sugoson (Uno, 20 Oct. 1950, A). HIROSHIMA. Taishaku-KYO (Na-ohara, 4 Sept. 1931, KYO); Mt. Azuma-yama (Kishino, 18 Aug. 1932, KYO); Hiroshmia-shi, Kabe-machi, NabaraKYO, alt. ca. 300m (Shimoda, 2 Oct. 1981, TI); Takada-gun, Mukai-hara, alt. ca. 300 m (Umebayashi, 13 Sept. 1977, KYO); Jinseki-gun, Toyomatsu-mura (Fu-kuoka & Kurosaki, 1968, KYO). TOTTORI. Tottori (Hiratsuka 12, TI); Iwami-gun, Kokufu-machi, Mt, Ohgino-sen (Tanaka, 27 Aug. 1978, KYO); Hino-gun, Nichinan-machi, nr. Abire, alt. ca. 600 m (Ohba & Akiyama 2153-2154, TI); Kedaka-gun, Shikano-machi, Mt, Washi-mine-sen (Tanaka, 5 Aug. 1975, KYO); Touhaku-gun, Misasa-mura, Ohse (Makino, 10 Aug., MAK); Touhaku-gun, Misasa-machi, Tawara, alt. 420 m (Ishizawa 17212, KYO); Mt. Daisen (Makino, Aug. 1906, MAK; Ikoma, 25 Aug. 1935, 9 Aug. 1936, KYO); Saihaku-gun, Daisen-cho, Mt. Daisen, near Daisen-ji Temple, alt. 1000 m (Terabayashi 378, KYO). SHIMANE. Nita-gun, Yokota-machi, near Nakamura, alt. ca. 400 m (Ohba & Akiyama 2131-2134, TI); Takobana (Okuyama & Utsumi 11626, TNS); Torikami-mura (Maruyama, Aug. 1937, TNS); Nishisusa (Maruyama, 3 Sept. 1944, TNS); Mt. Sanbe-san (Koidzumi, 26 July 1942, KYO); Hirata-tyo, Gakuenji, alt. 300 m (Murata, 27 Aug. 1958, KYO); Hirata-shi, Inome (Oka 39936, TNS); Matsue-shi, Tsuda-ku (Hayashi, 10 July 1936, KYO); Isl. Oki, Saigou-maclli, Araki (Oka 26020, TNS); Isls. Oki, Mt. Daimanji-san (Hori, 17 Aug. 1889, MAK); Isls. Oki, Shimamae, Kuroki-mura (Maruyama, 25 July 1937, KYO). YAMAGUCHI. Kuga-gun, Nishiki-machi, Usa (Oka 20598, TNS; 20600, KYO); Yoshiki-gun, Adzisu-machi (Shiomi s.n., KAG).

SHIKOKU. TOKUSHIMA. Myozai-gun, Ano-mura, Akawa, Hiroishi (Inobe, 15 July 1948, TNS); Myozai-gun, Ano-mura, Akawa (Inobe 681, TNS); TOKUSHIMA-shi, Nyuda-machi, Tsu-kinomiya (Takatou, 31 Aug. 1969, TNS); Oe-gun, Misato-mura, near Miyakura (Ohba & Akiyama 3764-3769, TI). EHIME. Onsen-gun, Nakajima-mura, Uwama (Yamamoto, 9 Sept. 1945, TI); Nishiuwa-gun, Iikata-mura, Kameura (Nomura, 16 Sept. 1953, KYO). KOUCHI. Mt. Chokou-san (unknown collector, 18 Oct. 1943, TI); Mt. Sasayama (Okuyama 16019, TNS); Takaoka-gun, Higashi-tunoyama-mura, Mt. Irazu-yama (Tagawa, 5 Sept. 1939, KYO); Nanokawa (Watanabe, Sept. 1888, A).

KYUSHU. In regionibus littoralibus secundum fretum v.o. Capellen (Pierot 872, L 908. 118-1056). FUKUOKA. Mt. Hiko-san (Yoshioka, 23 Sept. 1938, TOFO); Mt. Houman-zan (Nabeshima, 9 Oct. 1938, KYO); Buzen-shi, Mts. Inugatake, Osarebuchi-dani-Osao-touge (Hotta, 10 Aug. 1961, KYO); Asakura-gun, Kamiakidzuki-mura (Nakajima 26, TI). SAGA. Kashima-shi, Hiratani (Baba, 26 Aug. 1957, TNS); Oochi-machi, Yamase (Baba 19, KAG). NAGASAKI. (Oldham 330, L 908. 118-1055); Minamitakagi-gun, Hijikuro (Kunimi-machi, Hijikuro) (Makino, 10 Sept. 1906, MAK); Nagasaki, Hikoyama (Okada, 5 Oct. 1902, MAK); Mt. Tara-dake (Baba, 20 Aug. 1937, KYO); In vallibus montis ignivomi Wunzen (Pierot 312, L 908. 118-1077); Kitatakagi-gun, Oguri-mura (Yokoo, 7 Oct. 1938, TNS); Iki-gun, Gonoura-cho, Kamakifure-higashi, seaside (Mimoro 1710, KYO); Katsumoto-machi (Shinagawa, 24 Sept. 1955, TNS). Isl. Tsushima (Tashiro, 8 Aug. 1935, KYO); Kuroha-touge (Tashiro, 11 Aug. 1935, KYO); Shimoagata-gun, Uchiyama (Nakajima, 27 July 1933, KYO); Mt. Tara-san (Tashiro, 8 Aug. 1935, TNS); Shimoagata-gun, Izuhara-cho, on higher elevation of Mt. Aria-ke, alt. 550 m (Koyama 2658, TNS); Shimoagata-gun, Mt. Shiratake (Nakajima, 1 Aug. 1933, TI); Kuta-mura (Tashiro, 8 Aug. 1935, KYO); Shimoagata-gun, Himi, alt. 50m (Ohashi, Tateishi & Ohba67, 68, TI); Shimoagata-gun, Mizushima-cho, NW foot of Mt. Matsu-yama, alt. 100 m (Koyama 2753, TNS); Kamiagata-gun, WSW middle elevation of Mt. Mitake, alt. 200-400 m (Koyama 2837. TNS). KUMAMOTO. Mt. Sobo-san (Ohba, 15 Aug. 1934, KYO); Go-kanosho, alt. 1000 m (Hatusima & Sako 27122, KAG); Aso-gun, Yunotani (Sugino, 23 Oct. 1930, KYO); Aso-gun, Mt. Fukaba-yama (Yamashiro, 16 Aug. 1958, TNS); Namino (Momiyama, 7 Oct. 1958, TI); Aso-gun, Nojiriichi (Shimada 7848B, TNS); Aso-gun, Kugino-mura (Shi-mada 7857B & 16 July 1954, TNS); Aso, Mt. Neko-dake (Kôzuma, 25 Aug. 1926, TNS); Ashi-kita-gun, Sashiki (Kaneta 108, TI); Tenjin-touge (Yamashiro 4294, TNS); Aida (Mayebara 2111 & 31 July 1927, TI; 18 Sept. 1927, KYO); Watari (Mayebara, 28 Sept. 1924, KYO); Mt. Ichifusa-yama (Mayebara, 8 Oct. 1919, TYS); Kuma-gun, Kume-mura (unknown collector, 25 Aug. 1930, TI); Kamimura (Mayebara, 23 Aug. 1935, KYO); Amakusa-gun, Reihoku-cho, Tomioka (Tomiyama, Sept. 1968, KYO); Kikuchi-gun, Nishiharu-mura, alt. 400 m (Shimada, 15 Oct. 1961, KYO). OITA. Beppu-shi, Hotokezaki (Yamazaki, 28 Oct. 1928, KYO); Shimoge-gun, Tsutami-mura, Masaki (Yamazaki, 16 Sept. 1928, KYO); Mt. Kujû. (Makino in 1911, KYO). MIYAZAKI. Mt. Sobo-san (Tashiro, 31 Aug. 1933, KYO); Higashiusuki-gun, Toogoo-machi, Yamage-Oouchihara Dam (Murata 8585, TI); Mt. Souseki-san (Tashiro, 28 Aug. 1934, KYO). KAGOSHIMA. Aira-gun, Saishuyama-mura (Tashiro, 23 Sept. 1917, TNS); Mt. Kirishima, alt. 100-1000 m (Tashiro in 1917, A); Hayato-cho, alt, 200 m (Hatusima & Sako 25441, KAG); Aira-gun, Yoshimatsu, alt. 400 m (Hatusima & Sako 30273, KAG); Hioki-gun, Mt. Kinpou-san, at the foot (Hatusima 40548, KAG); Nishitara-mura (Doi, 26 Aug. 1932, KAG); Kaseda-shi, Toujinbaru (Ohba & Akiyama 2620-2626, TI).

Korea. CHOLLANAM. Isl. Heucksan Do (Daikokuzanto) (Ishidoya & Chung 3552, TI-Holotype of L. bicolor Turcz. sericea Nakai).

Amur (Komarov in 1895, LE).

Ussuri (Poretzky 122a, LE).

China. Mt. Po-hua-shan (Bretschneider 1877, LE). Manshuria Rossica. Bakuczan. In cozyletis montis (Komarov, 7 July 1895, TI); Manchuria Chinensis, prov. Murdenensy (Komarov, 16 July 1897, TI); Mandsuria austro-orientalis (Maximowicz in 1860, CAL); Coast of Manchuria (Wilford in 1859, CAL); Komoru-Ho, Hei-Ho (Takahashi 2344, TI); Hopeh in monte Hsiano-wu-tai-shan (Yabe, 28 July 1906, TI-Lectotype of L. ionocalyx Nakai); Liao-yang (Yabe, 15 Aug. 1909, TI); Lushan (Sato, 14 Aug. 1928, TI).

INNER MONGOLIA. Pailingmiao-Wuchuan (Togashi, 1 Sept. 1943, TI).

BEIJING. Pekin (Tatarinow in 1856, LE-Lectotype of L. bicolor Turcz. β. intermedia Maxim.); Peking, Nankou Pass (Wawura 986, LE). SHANDONG. (Maingay 35, CAL).

This species is distributed widely in Japan, but in the western and southern parts it grows at higher elevations. This species is distinguished from L. cyrtoborya by its wing being shorter than its keel-petal (Fig. 35). At Mt. Apoi, Hokkaido, however, a variety with a longer wing is found. It is thought that the relative length of the wing and the keel-petal is directly related to pollination by bees. This variety, therefore, is more or less isolated from typical ones reproductively. In Shikoku, especially the interior of Tokushima Prefecture and Ehime Prefecture, flowers with wings longer than the keel-petal as in L. cyrtobotrya are found. This Lesfedeza with these extra-ordinal flowers (see p. 18; Fig. 8) differs from var. nana by the taller habit and might be better treat as a local variety.

L. bicolor is reported from Taiwan (Huang & Ohashi, 1977), but we were unable to examine any specimens of L. bicolor from Taiwan. It is doubtful whether this species is distributed in Taiwan.

Forma niveoflora S. Akiyama et H. Ohba, nom. nov.

L. bicolor Turcz. f. albifiora Tatewaki in Trans. Sapporo Nat. Hist Soc. 16: 3 (1939), non Matsum. (1902)

Japanese name. Shirobana-yezo-yamahagi (Tatewaki, 1939).

Flores albi.

Distr. Japan (Hokkaido).

Specimens examined.

HOKKAIDO. TOKACHI. Ikeda-machi, Samamai (Okuda 4464, 25 Aug. 1937, SAPT-Holotype of L. bicolor f. albiflora Tatewaki); Ikeda (cult.) (Yokoyama, 25 Aug. 1937, SAPT).

L. bicolor f. albiflora Tatewaki is a later homonym of L. bicolor f. albiflora Matsu-mura (1902). Matsumura's albiflora is the white-flowered form of L. japonica, cv. Japonica, not of L. bicolor. This was previously known only from Hokkaido but it is expected to distribute sporadically in the range of this species.

Var. nana Nakai in Tennenkinenbutsu-oyobi-meisho-chosahokoku 12 (Veg, Apoi): 29 (1930)-Hara in Bot. Mag. Tokyo 49: 795 (1935).

Japanese name. Chabo-yamahagi (Nakai, 1930).

A perennial plant, 15-30 cm high. Terminal leaflet broadly elliptic to elliptic, 2-2.5 cm long, 1-2 cm wide. Raceme 1.5-2 cm long, shorter than subtending leaf. Calyx 3.5-4.5 mm long; lateral lobes triangular, acute, or more or less acuminate at the apex. The wing is nearly equal to or slightly longer than the keel-petal.

Distr. Japan (Hokkaido, Mt. Apoi).

Specimens examined.

HOKKAIDO. HIDAKA. On the upper part of Mt. Apoi (Nakai, Aug. 1928, TI-Holotype of L. bicolor Turcz. var. nana Nakai; Hara, 24 Aug. 1933, TI; Ohba & Akiyama, 22 Aug. 1984, no. 1-3, TI).

This variety grows only on the upper slopes of Mt. Apoi. At the foot of this moun tain and other localities in Hokkaido var. bicolor is abundant. Some var. nana have a calyx with an acuminate apex and short inflorescence, and are very similar to L. cyrto-botrya. But var. nana differs from that in the texture of leaf, the shape of fruit, and some other vegetative features.