Report on the poriferan specimens deposited in the Department of Zoology, the University Museum, the University of Tokyo - (1) Hexactinellida
Kazunari Ogawa1), Wataru Abe2) and Rei Ueshima2, 3)
1)Z. Nakai Laboratory, 2-53-11, Koenjiminami, Suginami-ku, Tokyo, 166-0003 Japan
2)Department of Biological Sciences, Graduate School of Science, University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo, 113-0033 Japan
3)Corresponding author: Reprint request to rueshima@biol.s.u-tokyo.ac.jp
Summary
The hexactinellid poriferan collection deposited in the Department of Zoology, the University Museum, the University of Tokyo, consists of 315 lots of dry specimens, 470 lots of wet specimens and 28 lots of fossil specimens. The hexactinellid specimens are voucher specimens studied by Isao Ijima and Yaichiro Okada of The Tokyo Imperial University. Most of the hexactinellid specimens were collected from Sagai Bay about 100 years ago but the collection also includes some specimens collected by Siboga's and Albatross's expeditions.
The collection includes following type specimens of the hexactinellid taxa described by I. Ijima and Y. Okada: Aphrocallistes beatrix orientalis Ijima 1916 [syntype], Aulocystis zitteli sibogae Ijima 1927 [syntype], Aulosaccus mitsukurii Ijima 1898 [holotype], Calyptorete ijimai Okada 1925 [paralectotype], Chalaronema sibogae Ijima 1927 [paratype], Chaunoplectella cavernosa Ijima 1896 [holotype], Chaunoplectella stelleta Ijima 1927 [holotype], Crateromorpha meyeri var. rugosa Ijima 1898 [syntype], Crateromorpha meyeri var. tuberosa Ijima 1898 [syntype], Euplectella curvistellata Ijima 1901 [holotype], Euplectella timorensis Ijima 1927 [holotype and paratype], Eurete trachydocus Ijima 1927 [holotype], Farrea hanitschi Ijima 1927 [holotype], Hexactinella lingua Ijima 1927 [holotype and paratype], Hyalascus similis Ijima 1904 [holotype], Hyalonema (Coscinonema) kirkpatricki Ijima 1927 [holotype and paratype], Hyalonema (Cyliconema) keiense Ijima 1927 [syntype], Hyalonema (Cyliconema) timorense Ijima 1927 [holotype], Hyalonema (Euhyalonema) intermedium Ijima 1927 [holotype], Hyalonema (Euhyalonema) keianum Ijima 1927 [holotype and paratype], Hyalonema (Paradisconema) vosmaeri Ijima 1927 [holotype], Hyalonema (Pteronema) topsenti Ijima 1927 [holotype and paratype], Hyalonema (Thamnonema) thamnophorum Ijima 1927 [syntype], Hyalonema ovatum Ijima 1895 [syntype?], Hyalonema owstoni Ijima 1894 [syntype], Leucopsacus orthodocus Ijima 1898 [lectotype], Lonchiphora inversa Ijima 1927 [syntype], Lophocalyx suluanus Ijima 1927 [holotype and paratype], Pararete farreopsis fragiferum Ijima 1927 [holotype and paratype], Pararete farreopsis subglosum Ijima 1927 [holotype and paratype], Pheronema pepo Ijima and Okada 1938 [holotype], Pheronema weberi Ijima 1927 [syntype], Placosoma paradictyum Ijima 1903 [holotype], Regadrella komeyamai Ijima 1901 [syntype], Rhabdocalyptus capillatus Ijima 1897 [syntype], Rhabdocalyptus unguiculatus Ijima 1904 [holotype and paratype], Semperella similis Ijima 1927 [holotype], Staurocalyptus affinis Ijima 1904 [syntype], Staurocalyptus entacanthus Ijima 1904 [holotype], Staurocalyptus glaber Ijima 1897 [syntype], Staurocalyptus heteractinus Ijima 1897 [holotype], Staurocalyptus pleorhaphides Ijima 1897 [syntype], Sympagella anomala Ijima 1903 [syntype], Tretopleura candelabrum Ijima 1927 [holotype and paratype], Tretorete incertum Ijima 1927 [syntype], Vitrollula fertile Ijima 1898 [syntype], Vitrollula namiyei Ijima 1898 [syntype].
In the following taxa, type specimens are deposited in some foreign museums but a fragment of each type exists in our collection: Acanthascus pachyderma Okada 1932 [a fragment of holotype], Aphrocallistes aleutiana Okada 1932 [a fragment of syntype], Aphrocallistes intermedia Okada 1932 [a fragment of syntype], Aphrocallistes yatsui Okada 1932 [a fragment of holotype], Aulosaccus albatrossi Okada 1932 [a fragment of holotype and paratype], Aulosaccus fissuratus Okada 1932 [a fragment of paratype], Aulosaccus fissuratus shimushirensis Okada 1932 [a fragment of holotype], Aulosaccus pinularis Okada 1932 [a fragment of holotype], Aulosaccus solaster Okada 1932 [a fragment of holotype], Aulosaccus tuberculatus Okada 1932 [a fragment of syntype], Eurete irregularis Okada 1932 [a fragment of syntype], Eurete nipponica Okada 1932 [a fragment of holotype], Eurete sacculiformis Okada 1932 [a fragment of syntype], Farrea kurilensis Okada 1932 [a fragment of holotype], Farrea sollasii yakushimensis Okada 1932 [a fragment of holotype or paratype], Farrea watasei Okada 1932 [a fragment of holotype], Hyalascus attenuatus Okada 1932 [a fragment of holotype], Hyalascus sagamiensis Ijima 1896 [a fragment of holotype], Hyalonema (Coscinonema) kirkpatricki globosum Okada 1932 [a fragment of paratype], Hyalonema (Coscinonema) ovatum Okada 1932 [a fragment of holotype], Hyalonema (Cyliconema) apertum solidum Okada 1932 [a fragment of holotype and paratype], Hyalonema (Cyliconema) hozawai Okada 1932 [a fragment of holotype and paratype], Pheronema globosum kagoshimensis Okada 1932 [a fragment of holotype], Pheronema ijimai Okada 1932 [a fragment of holotype and paratype], Pheronema surugensis Okada 1932 [a fragment of holotype], Rhabdocalyptus bidentatus Okada 1932 [a fragment of holotype], Rhabdocalyptus heteraster Okada 1932 [a fragment of holotype or paratype], Staurocalyptus rugocruciatus Okada 1932 [a fragment of holotype].
Type specimens of the following taxa are lost: Anomochone expansa Ijima 1927, Anomochone globosa Ijima 1927, Bathydorus pedunculatus Ijima 1927, Bolosoma cavum Ijima 1927, Chaunoplectella spinifera Ijima 1903, Crateromorpha pachyactina Ijima 1898, Diapleura maasi Ijima 1927, Eurete freelandi Ijima 1927, Eurete schmidti kampeni Ijima 1927, Eurete schmidti treubi Ijima 1927, Farrea beringiana Okada 1932, Farrea lendenfeldi Ijima 1927, Farrea nodulosa Ijima 1927, Farrea occa cuspidata Ijima 1927, Farrea occa mammillata Ijima 1927, Farrea occa subclavigera Ijima 1927, Farrea spirifera Ijima 1927, Hexactinella rugosa Ijima 1927, Hexactinella vermiculosa Ijima 1927, Hexactinella? spongiosa Ijima 1927, Hyalonema (Corynonema) rotundum Ijima 1927, Hyalonema (Leptonema) flagelliferum Ijima 1927, Hyalonema pellucidum Ijima 1894, Leucopsacus scoliodocus Ijima 1898, Myliusia verrucosa Ijima 1927, Pararete baliense Ijima 1927, Pararete farreopsis jakosalemi Ijima 1927, Pararete freeri Ijima 1927, Pararete kangeanganum Ijima 1927, Pararete semperi Ijima 1927, Pheronema placoideum Ijima and Okada, 1938, Pleurochorium cornutum Ijima 1927, Priphragella irregularis Ijima 1927, Priphragella parva Ijima 1927, Psilocalyx wilsoni Ijima 1927, Regadrella cylindrica Ijima 1927, Rhabdodictyum kurense Ijima 1927, Rossella longispina Ijima 1896, Sclerothamnopsis schulzei Ijima 1927, Staurocalyptus celebesianus Ijima 1927, Staurocalyptus microchetus Ijima 1898, Staurocalyptus tubulosa Ijima 1904, Tretodictyum pumicosum Ijima 1927, Tretodictyum schrammeni Ijima 1927.
The following names found on some labels of the collection are Ijima's manuscript names: Acanthascus tubus, Farrea awokii, Farrea clavata, Hyalonema claviferum non Hyalonema (Corynonema) clavigerum (Schulze, 1886), Hexactinella polystomata, Regadrella tubus, Rhabdocalyptus maximus, Rhabdocalyptus spinifer non Chaunoplectella spinifera Ijima, 1903, Staurocalyptus japonicus non Staurocalyptus japonicus in Ijima (1903), Staurocalyptu crassifibratus. Some of the specimens with such manuscript names may be candidates for new species.
Hyalonema ovatum Ijima, 1895 and Hyalonema (Coscinonema) ovatum Okada,1932 are primary homonym. Here we propose a new name, Hyalonema (Coscinonema) okadai Ogawa and Ueshima (nom. nov.), as a replacement name for Hyalonema (Coscinonema) ovatum Okada,1932.
