Lespedeza Buergeri Miquel

In Ann. Mus. Lugd.-Bat. 3: 47 (1867) et Prol. Fl. Jap., 235 (1867)-Maxim. in Acta Hort. Petrop. 2: 353 (1873), excl. quoad specim. Wilford (cf. Schindler, 1913), Oldham 333-Franchet & Savatier, Enum. Pl. Jap. 1: 101 (1875)-Forbes & Hemsley in Journ. Linn. Soc. Bot. 23: 179 (1887), pro parte-Diels in Engl., Bot. Jahrb. 29: 415 (1901)-Matsumura in Bot. Mag. Tokyo 16: 70 (1902); Ind. Pl. Jap. 2: 267 (1912)-Schneider, Ill. Handb. Laubholz. 2: 113, fig. 70k. (1907)-Pampanini in Nuov. Giorn. Bot. Ital. n. ser. 17: 398 (1910); 18: 123 (1911)-Schindler in Engl., Bot. Jahrb. 49: 580 (1913); in Sargent, Pl. Wils. 2: 106 (1914)-Makino & Nemoto, Fl. Jap., 734 (1925)-Rehder in Journ. Arn, Arb. 7: 171 (1926)-Momiyama in Journ. Jap. Bot. 9: 127 (1933)-Ohwi, Fl. Jap., 789 (1953); Fl. Jap. rev. ed., 678 (1965); Fl. Jap. Eng. ed., 559 (1965); Fl. Jap. new ed., 789 (1975)-Fu in Acad. Sin. Bot., Ill. Important Chin. Pl. Legumimsae, 521, fig. 514 (1955)-Kitamura & Murata, Col. Ill. Herb. Pl. Jap. 2: 98 (1961)-Kung in Chinese Journ. Bot. 1: 20 (1963)-Hatusima in Mem. Fac. Agr. Kagoshima Univ. 6: 6 (1967), excl. ssp. praecox (Nakai) Hatusima, ssp. tricolor (Nakai) Hatusima-Acad. Sin. Bot., Iconogr. Cormophyt. Sin. 2: 460, fig. 2649 (1972)-Ohashi in Satake et al., Wild Fl. Jap. Herb. 2: 205 (1982). [Plates 21 & 22; Fig. 46]

L. Oldhamii Miq. in Ann. Mus. Bot. Lugd.-Bat. 3: 48 (1867) et Prol. Fl. Jap., 236 (1867).

L. Sieboldii Miq. in Ann. Mus. Bot. Lugd.-Bat. 3: 47 (1867) et Prol. Fl. Jap., 235 (1867), pro parte, quoad specim., Textor.

L. Buergeri Miq. var. Oldhamii (Miq.) Maxim. in Acta Hort. Petrop. 2: 354 (1873)-Matsumura, Ind. Pl. Jap. 2: 267 (1912).

L. bicolor Turcz. β. intermedia Maxim. in Acta. Hort. Petrop. 2: 356 (1873), pro parte.

L. Buergen Miq. f. longiracemosa H. Koidzumi in Journ. Pl. Iwateken 2: 77 (1937), typum non vidi.

L. Buergeri Miq. f. subbiloba H. Koidzumi in Journ. Pl. Iwateken 2: 77 (1937), typum non vidi.

L. Buergeri Miq. var. tetraloba H. Koidzumi in Journ. Pl. Iwateken 2: 76 (1937), typum non vidi.

L. Buergeri Miq. f. Oldhamii (Miq.) Sigimoto, New Keys Jap. Trees, 463 (1961), Japanese name, Kihagi (Matsumura, 1912).

A perennial plant, 1-3 m high. Stems ascending, 0.5-1 m high, 1-5 cm in diameter; branched in upper parts (a few branches coming from basal part of the stem). Branches terete, spreading or sometimes dependent later; when young slightly angular with ascending appressed-sericeous or ascending hairs (hairs 0.3-0.6 mm long, yellowish), later glabrescent; most distal parts die in winter season, proximal parts (and stem) with axillary and adventitious winter buds (winter buds at lower parts mostly dormant).

Leaves trifoliolate, petiolate, stipulate, alternately arranged (phyllotaxy 1/2). Stipules free, linear-triangular to linear, 2-7 mm long, brownish, persistent. Petioles 0.5-4 cm long, similar to the young branch. Rachides 5-15 mm long, similar to the petiole. Terminal leaflets (at middle parts of branches) petiolulate; petiolules 1-2 mm long, swollen; lamina 1-5 cm long, 0.5-3 cm wide, entire, (narrowly to broadly) ovate or elliptic, round or cuneate at the base, acute or obtuse at the apex (the apex itself with or without a point), upper surface glabrescent (except along the midrib) or sericeous (hairs ca. 0.5 mm long) at anthesis, lower surface appressed-sericeous (hairs ca. 0.5 mm long, yellowish); lateral ones similar to terminal but somewhat smaller.

Inflorescence axillary racemous, one (rarely two to three) per axil, 2-4(-7) cm long including the peduncle, somewhat compactly to loosely 4-10-flowered; peduncles 0.5-1 cm long, hairy like the young branch. Primary bracts ovate, about 1 mm long, pubescent, persistent; secondary bracts similar to primary ones.

Flowers 8.5-10.3 mm long at anthesis; pedicels 0.8-2 mm long terete, curved, tomentose. Bracteoles at the base of the calyx round to elliptic, 1-1.5 mm long, pubescent outside, glabrous inside, persistent. Calyx 1.8-2.9 mm long, campanulate, four-lobed near the middle part, appressed pubescent; tube 1.1-1.4 mm long; lobes subequal in length or the lower one longest; lateral ones 0.7-1.7 mm long, 0.7-0.9 mm wide, broadly ovate or triangular-lanceolate, obtuse or acute at the apex but not acuminate; upper one slightly two-clefted.

Standard shorter than keel-petal and longer than (or sometimes as long as or shorter than) wings, keel-petal longer than wings (K>S≥W). Standard pale yellow with purple patches near the base inside, whitish outside, 7.3-8.8 mm long with distinct claw, 5.0-6.0 mm wide; lamina round to elliptic or broadly obovate, round or retuse at the apex with or without a point, inflexed-auriculate near the base, spreading in anthesis from the part near the base; auricles well developed; claw whitish, 1.1-1.5 mm long. Wings purple or deep red-purple, 7.3-8.6 mm long with distinct claw; lamina narrowly obovate to oblong, 5.0-6.4 mm long, 2.0-2.5 mm wide, auriculate at upper basal part, slightly or minutely auriculate or tapering at the lower basal part; claw 2.2-2.7 mm long, whitish. Keel-petal 8.0-10.1 mm long with distinct claw; amina obovate, 5.7-7.7 mm long, 2.7-3.5 mm wide, pale yellow; claw 2.4-2.8 mm long, whitish. Stamens 10, nearly the same length, 7.8-9.9 mm long, diadelphous. Anthers elliptic shallowly, retuse at the apex, ca. 0.5 mm long; before anthesis yellow. Pistils 7.9-10.0 mm long; ovary narrowly elliptic, ca. 1.8 mm long, ca. 0.5 mm wide at anthesis, pubescent; style 6-8 mm long, pubescent at the basal part, glabrescent at the apical part, stalked; stalk ca. 1.2 mmlong.

Fruits compressed, narrowly elliptic, 11-15 mm long, 5-5.5 mm wide, stalked; stalk 1.5-2 mm long, slightly to densely pubescent or nearly glabrous.

Seeds reniform, 3.5-4 mm long, 2-2.5 mm wide.

Distr. Japan (Hoshu, Shikoku, Kyushu), E. China.

Voucher and representative specimens.

Japan (Buerger s.n., GH; L 951326 132); (Textor s.n., L 908.118-2364-Syntype of L. Sieboldii Miq.); (Oldham 335, L 908.118-1398-Holotype of L. Oldhamii Miq., ut "Japan & Korea Arch. coll."); ad littus maris. . . (Siebold s.n., L 908.118-1368-Syntype of L. Buergeri Miq.).

HONSHU. IWATE. Kuji-shi, Ookawame (Murata & Tabata, 28 July 1967, KYO); Ryusenkutsu (Shimizu, 1 Aug. 1956, KYO); Shimohei-gun, Iwaizumi-cho, Mt. Ureira (Shimizu, 11 Aug. 1956, KYO); Shimohei-gun, Tanohata-mura, Naraiga (Kikuchi, 28 Sept. 1967, TUSG); Shimohei-gun, Kariya-mura, Oshikado-touge, alt. 300-400 m (Kanai, 2 Sept. 1959, TI); Shimohei-gun, Akka-mura (Hosoi, 15 Sept. 1952, A); Kamihei-gun, Oodzuchi-machi, Kirikiri (Kanai, 3 Sept. 1959, TI); Kamihei-gun, Oohashi (Toba 15, TI); Miyako-shi, Omoe (Yamamoto, 28 July 1971, TNS); Kesen-gun, Takada (Toba, 538, TI). MIYAGI. Jogi-onsen (Kamio, Aug. 1934, Kyo); Mt. Kurikoma (Okuyama, 21 Aug. 1952, TNS); Ojika-gun, Onagawa-cho, Urajuku (Kikuchi, 21 Oct. 1967, TNS); Mono-gun, Naruse-cho, Miyato-jima (Naito, 19 Aug. 1973, TUSG); Daitodake (Ôhashi, 2 Sept. 1960, TUSG); Kawasaki, Kamafusa, along River Goishi, alt. 150m (Sohma 1237, TUSG); Zaou, Aone (Satake & Ito, 17 Oct. 1958, TNS); Tougatta, Torii (Momiyama, 12 July 1936, TI); Sendai, Idohama (Kikuchi & Okazaki, 17 Oct. 1969, TUSG); Sendai, Aobayama (Kimura, Sugaya, Saito & Mori, 10 July 1959, TI). YAMAGATA. Yamaike (Doi in 1966, TNS). FUKUSHIMA. Date-gun, Kunimi-machi, Kaida (Sato, Kaneko & Hayashi, 3 Aug. 1965, TUSG); Shiogama (Jack, 27 Aug. 1905, GH). GUNMA. Adzuma-gun, Adzuma-machi, Haramachi (Masuda in 1940, MAK); Myogisan (Ishii 8, TNS)*; Tano-gun, Nakazako-mura, Kamigahara, Mt. Kanoosan, alt. 1100m (Murata, Ohba, Kadota & Midorikawa 1072, TI). TOCHIGI. Shiobara, Shiogama-Shionoyu (Kanai, 27 May 1959, TI); Nikko (Matsumura, 29 July 1877, TI); Nikko-shi, Jakkou-no-taki (Ohba & Akiyama 1383 & 1384, TI); Mt. Akanagi-yama (Nakai, 5 Aug. 1931, TI). IBAEAKI. Tsukuba (Oowatari, 9 July 1895, TI); Nishiibaraki-gun, Minamikawane-mura (Iwama-mura) (Okamoto, 4 Aug. 1904, MAK); Tsuchiura-shi, Hirookahara, Otto-numa (Makino, 15 Aug. 1913, MAK). CHIBA. Awakominato-machi, Mt. Uchiurayama, alt. less than 300 m (Ohba et al. 6808042, TI); Mt. Kiyosumi-yama (Kiyozumi-yama) (Okuyama 22724, TNS); Kimitsu-gun, (Akimoto-mura), Mt. Kano-san, alt. 150m (Hurusawa 109, TI); Mt. Mitsuishi (Okuyama 22877, TNS); Okitsu-machi (Furuse, 16 Oct. 1954, TNS). SAITAMA. Chichibu, Takashino, Sadamine (Moriya, 30 Sept. 1961, TNS)*; Chichibu-gun, Nogami-machi, Nagatoro (Murata, Tateishi, &Kadota 743,TI); Chichibu, Kagemori (Moriya, 7 Aug. 1961, TNS)*; Chichibu, Mt. Mitsumine-san (Sakurai, July 1892, TNS)*; Chichibu, Mt. Kumakura-yama (Moriya, 5 Sept. 1970, TNS); Chichibu-gun, Ryoogami-mura, Sakamoto, Mt. Futagoyama, alt. 1000m (Murata, Tateishi & Midorikawa 2441, TUSG); Kamabuse-touge (Setsuda, 13 Oct. 1940, TNS)*; Hannou, Mt. Tenran-zan (Hattori, 14 May 1922, TI); Hanno-shi, Mt. Izugatake (Akiyama 338, TI). TOKYO. Mt. Tenso-zan (Takeuchi, 10 Aug. 1948, TI); Kawanoriyama (Kimura, Sept. 1951, TI); Okutama, Hatonosu (Kanai 720, TI); Mt. Takamnizu-yama (Mt. Takamizu) (Mizushima, 4 Sept. 1954, TI); Mt. Gozen-yama (Mizushima, 7 July 1946, TI); Nishitama-gun, Hinohara-mura, Fukamata, alt. ca. 700m (Ohba 1033, TUSG); Itsukaichi, Sengen-one (Ohba & Akiyama s.n., TI); Itsukaichi, Tokisaka (Akiyama 294, 297, TI); Takao-san, alt. 300 m (Wilson in 1914, A); Mt. Jinba-yama (Kazami, 14 Oct. 1962, TNS); Asakawa, Koshi-tasawa (Kanai 705, TI); Sekido (Mizushima 2493, TI). KANAGAWA. Yokohama (Hisauchi 409, TI); Kamakura (Momiyama 682, TI); Zushi-shi, Zinmu-ji (Ohba & Murata 669, TI); Kurihama (Makino in 1934, MAK); Miura-kaigan (Ito, 12 Nov. 1970, TI); O-iso (Momiyama 944, TI); Mt. Tanzawa-yama, Mt. Yake-yama (Yamazaki, 6 Oct. 1942, TI); Aikou-gun, Susu-gaya-mura, Teraie-Monomi-touge, alt. 400m (Kanai, 10 Nov. 1957, TI); Ashigarakami-gun, Kamihatano-mura, Horinishi (Kanai, 27 Aug. 1956, TI); Yamakita (Komori 1975, TI); Ashigarakami-gun, Mt. Takamatsu-yama (Henmi 2304, TNS); Hakone, Mt. Kintoki-yama (Mt. Kintoki) (Momiyama, 7 Sept. 1951, TI); Hakone (Maximowicz in 1862, GH); Ashiga-rashimo-gun, Hakone-machi, Tounosawa (Ohba & Akiyama 614-615, TI); Manadzuru (Henmi, 20 July 1952, TNS). YAMANASHI. Yanakawa-Mt. Kuratake-yama (Mizushima, 26 June 1949, TI); Mt. Mitsutouge-yama (Sawada, 23 Aug. 1930, TI); Mts, Misaka, Mitsuke Pass (Kanai, 18 Sept. 1955, TI); Minamitsuru-gun, Funatsu-mura, on shore of Kawaguchi Lake, alt. 830m (Furuse, 11 Sept. 1958, A); Nisiyatusiro-gun, Lake Shoji-ko (Kanai, 15 Sept. 1957, TI); Nisiyatusiro-gun, Kamikuisshiki-mura, Onnasaka-Koseki (Kanai, 15 Sept. 1957, TI); Nisiyatusiro-gun, Ookawachi-mura, Shionosawa (Kanai, 22 Sept. 1957, TI); Mt. Misho-tai-yama, Shimosha-Okusha (Kanai, 3 Nov. 1956, TI); at the foot of Mt. Fuji-san, Lake Motosu-ko, Ryugatake rindou (Kanai, 1 June 1958, TI); Mt. Fuji-san, Aoki-ga-hara, near Hyoketsu (Ohba 6707, TUSG); Mt. Minobu-san (Okuyama, 26 Sept. 1954, TNS); Kobuchi-zawa (Togashi 7390, TI); Nirasaki-shi, Anayama, alt. 550 m (Tateishi 4486, TI); Mt. Houou-zan, alt. 1300m (Uematsu, 1 Oct. 1949, TI); Mt. Koma-ga-take (Satake 1556, TI); Mts. Tenshi, Okunoyu (Shimobe) (Kanai, 21, Aug. 1955, TI). NAGANO. (Tschonoaki in 1864, GH); Karuizawa, Myozenji (Greatrex K25/30, TI); Minamisaku-gun, Nakagomi-machi, Arafune (Sato, 5 Aug. 1958, TI); Komoro, Mt. Nunobikiyama (Hasegawa, 12 July 1964, TI); Minamisaku-gun, Minamiaiki-mura, Sankawa-Mt. Sekibutsu, alt.140 0-1920 m (Murata & Ohba 5239, TI); Kawanakajima (Matsumura, 16 July 1884, TI); Chiisagata-gun, Shioda-cho, Besshoonsen (Kitagawa 5398, KYO); Chiisagata-gun, Niuchi-mura (Momose, 8 Aug. 1931, TI); Mt. Hachibuse (Momose 664, TI); at the foot of Mt. Senjo-dake, Todai (Uematsu, 24 Sept. 1949, TI); Ontakesan (Koidzumi, Aug. 1910, TI); Mts. Akashi, Mt. Hijiri-dake, Nishizawa, alt. 1400 m (Yamazaki, Matsuda & Fujita, 6 Aug. 1954, TI); Nakasendou, Agematsu-Fukushima (Ito, 15 Aug. 1891, TNS); Kamiina-gun, Inasato-mura, Mitsumine-gawa, alt. 1000 m (Matsuda, 16 July 1954, TI); Shimoina-gun, Ueura (Koidzumi, 23 Sept. 1932, TNS); Shimoina-gun, Aiji-mura, Komaba-Sonohara (Kanai, 29 July 1957, TI); Shimoina-gun, Hiraoka-machi (Takei, 12 Aug. 1957, TNS); Shimoina-gun, Ooshikamura, Ochiai (Asano 9239, TI); Shimoina-gun, lida-shi, Kazekoshi (Furusawa, Aug. 1940, TI); Shimoina-gun, Yamabukimura, Shimo-machi (Asano 11229, TI); Shimoina-gun, Minamiwadamura, Sokoike-Nagoyama (Asano 8120, TI); Shimoina-gun, Hiraokamura, Orochi (Asano 24425, TI). SHIZUOKA. Numadzu (Tako, 15 Aug. 1907, MAK); Mts. Ashitaka, Takayama (Kanai 6078, TI); Fuji, Subashiri-guchi (Muramatsu, 30 July 1924, TI); Fuji-gun, Kamiide-mura, Inokashira-Wakimizu-touge, alt. 1000 m (Kanai, 23 Sept. 1957, TI); Fuji-gun, Kamiide-mura, Mt. Kenashi-yama, alt. 1100 m (Kanai, 18 May 1958, TI); Ookouchi-mura, Mafuji-mura (Furuse 19885, TI); lhara-gun, Mt. Kunou-zan (Mizushima 1640, TI); Umegashima, ûtaki (Okuyama, 3 Oct. 1951, TNS); Abe-gun, Ikawa-mura, Tashiro-gawa (Sugimoto 20435, TI); Haibara-gun, Honkawame-cho, the top of Mt. Fudou-the foot of Mt. Morosawa, alt. ca. 1000 m (Koyama, Miki & Yahara 134, KYO); Shida-gun, Setonotani-mura (Shimizu 157, TI); Iwata-gun, Fukuroi-machi (Hashimoto, 24 July 1932, TNS); Kakegawa-shi, the foot of Mt. Ogasayama (Ohba 3496, TI). AICHI. Tomiyama-mura, Kawauchi (Torii 12404, TNS); Kita-shitara-gun, Tsugu-mura (Torii, 18 July 1954, TUSG); Kitashitara-gun, Miwa-mura, Uren-gawa (Yamazaki, 30 July 1957, TI); Minamisliitara-gun, Hourai-machi, Mutsudaira, alt. 100-150 m (Mimoro, Tsugaru & Nishiyama 2004, TNS); Nagashino-mura (Torii 6108, TUSG); Shinshiro-shi (Torii 6135, TUSG); Mt. Ishimaki-yama (Torii 6162, TUSG); Moya (Makino, 30 Sept. 1905, MAK); Toyohashi, Ishimaki, Wada (Torii, 20 Aug. 1967, KYO). GIFU. Mt. Ena-san (Sakurai, July 1903, TNS); Toki-gun, Tajimi (Shiota, 17 July 1933, KYO); Toki-gun, Tsumald-machi (Shiota, 17 Sept. 1933, KYO); Toki-gun, Ichinokura-mura (Shiota, 15 Aug. 1929, KYO); Kamo-gun, Higashi-shirakawa-mura (Shiota 8096, A); Mugi-gun, Suhara-mura (Shiota 6487, A); Mugi-gun, Mino-machi (Shiota 5390, A); Mugi-gun, Itadori-mura, NW foot of Mt. Kooka-san, Kabe-Taragadani-goe (Fukuoka & Yamashita 222, KYO); Kani-gun, Toyooka-machi, Ueno (Ima-dzue, 21 Aug. 1933, A); Gujo-gun, Yatomi-mura (Shiota, 27 July 1932, TI); Gujo-gun, Hachi-man-machi (Hiroe, 22 Aug. 1948, KYO); Gujo-gun, Kawai-mura (Shiota 7654, A); Gujo-gun, Yatomi-mura, Banba (Shiota 7253, A); Ishidzu-mura (Sugino, 20 Aug. 1952, TNS); Yourou-gun, Yourou-machi, Yourou-no-taki (Kanai, 1 Oct. 1957, TI). SHIGA. Mt. Ibuki-yama (Matsumura, 1 Aug. 1881, TI); Gamou-gun, Nishiooji (Hashimoto, 22 Aug. 1947, TNS); Sugi (Hashimoto, 8 Aug. 1945, TNS); Mt. Watamuki-yama (Senbongi-Kumano) (Araki 5744, SHO); Koga-gun, Tsuchiyama-cho, Tamura-jinja, alt. 260m (Murata, 24 Oct. 1964, KYO); Mt. Hira-san, Kido (Hashimoto, 7 Nov. 1944, SHO); Katatatyo, Tochu-Sakashiba (Kitamura & Murata, 5 Nov. 1961, KYO); Ootsu-shi, Mt. Ousaka-yama (Hashimoto 11347, KYO). MIE. Komono, Yunoyama (Sono, 23 Aug. 1908, TNS); Mie-gun, NE of Mt. Gozaisho, Yuno-yama-Asake, alt. 600-800 m (Fukuoka, 4 Aug. 1963, KYO); Inabe-gun, Ishigure-inura, Mt. Ryuêgadake (Fukuoka, 6 Aug. 1963, KYO); Suzuka-gun, at the foot of Mt. Nonobori, around so-called Byôbuiwa (Shimizu, 16 July 1959, KYO); Aki-gun, Mt, Kyogamine (Imai, 10 Oct. 1976, SHO); Ise Shrine forest, Kamijiyama (Nitta, 12-16 Aug. 1963, KYO); Watarai-gun, Mise-tani (Oomiya-machi) (Umemura, 11 Aug. 1895, MAK); Watarai-gun, Shimadzu-mura, Kowa-mura (Kuwana, 20 July 1905, MAK); Mt. Asama-yama (Momiyama 1037, TI); Ayama-gun, Ueno (Koreishi, July 1903, MAK) ; Isshi-gun, Tarou-mura, Mt. Kuroso (Kuruson) (Kanai, 20 Sept. 1962, TI); Naka-gun, Akame (Tagawa, 14 Aug. 1950, KYO); Oaugidani Hinode-dake-Momonoki, alt. 600m (Yamazaki, Yamazaki & Morita, 6 June 1973, TI); Minamimuro-gun, Gogou-mura, Momozaki-Yuya (Kanai, 28 April 1957, TI); Minamimuro-gun, Mihama-cho, Huden-toge, alt. 250 m (Mimoro, Tsugaru & Deguchi 4285, KYO) ; Ootou-yama (Okamoto, 27 Oct. 1952, TNS); Mt. Odaigahara (Ohwi 9103, TNS). NAKA, Oodai-ga-hara, Shinonoha (Ohwi, 3 Sept. 1936, KYO); Yoshino-gun, Kawakami-mura, Yamato (Koidzumi, Sept. 1924, TNS); Yoshino-gun, Tenkawa-mura, Mt. Ohmine, Mt. Chosendake-Tu-bonouchi, alt. ca. 700m (Fukuoka & Hotta, 6 Aug. 1962, KYO); Yoshino-gun, Shimokita-yama-mura, Zenki-Zenkiguchi (Murata & Shimizu, 18 July 1954, KYO). WAKAYAMA. Kumano-shi, Isato-machi, Ooidani, Kitayama River, upstream (Furuse, 13 Oct. 1961, A); Higa-shimuro-gun (ut "Kitamuro-gun"), Kitayama-mura, Komatsu (Furuse, 6 Oct. 1962, A); Higashimuro-gun, Kumanogawa-cho, Koguchi-Ohara, along the River Wadagawa (Murata, 15 Oct. 1960 & 13587, TI); Mt. Nachi-san (Matsumura, 29 July 1883, TI); Shinguu (Chihomine) (Satake & Okuyama, 11 Oct. 1942, TNS); Higashimuro-gun, Takada-mura (Ito, 5 Oct. 1930, TI); Higashimuro-gun, Hongu-cho, Mt. Ooto-zan, interior of Shizukawa, Oosugi-dani (Naruhashi 1759, KYO); Higashimuro-gun, Koza-cho (Uno, 21 July 1952, A); Isl. Ooshima (Tsukamoto, 9 Oct. 1940, KYO); Nishimuro-gun, Nakaheji-cho, Fukusada-Nonaka (Mimoro, Tsugaru & Fujimoto 4062, KYO); Nishimuro-gun, Tanabe-machi (Nakajima 1310, TI); Namera-dani (Araki 5742, SHO). KYOTO. Sokaku-gun, Kasagi-cho, Kasagiyama, alt. ca. 150m (Murata 20457, KYO); Kyoto-shi, Ukyo-ku, Mt. Atago-yama, alt. 100m (Murata 20445, KYO); Hodzu-kyo (Murata 5950, TNS). HYOGO. Takarazuka-shi, Horai-kyo (Togashi, 23 June 1968, KYO); Miki-shi, Shimizu-cho, Mitsuda, Yamadagawa, alt. 100-200 m (Murata & Nishimura 201, KYO); Kobe-shi, Kita-ku, north slope of Mt. Rokkô, Gokuraku-chaya-Arima-cho, alt. 400-800 m (Shimizu & Kurosaki 14, SHO); Shikama-gun, Yumesaki-cho, Koura (Kawara)-keikoku (Hashimoto 10061, SHO); Isl. Awaji, Inohana-dani (Tashiro, 23 Sept. 1927, KYO). OKAYAMA. Wake-gun, Saeki-machi, Ohara (Ookubo, 19 Aug. 1977, KYO)*; Atetsu-gun, Arato-mura, Kounoie (unknown collector, 1 July 1929, KYO); Akaiwa-gun, Takatsuki-mura, Mt. Takakura (Uno, 15 Aug. 1952, A).

SHIKOKU. KAGAWA. Shoozu-gun (Shoodo-shima), Kankakei, alt. 700 m (Ohashi, Ohba & Murata, 10 Nov. 1975, TI); Shouzu-gun, Kusakabe-mura (Honma, 5 Aug. 1905, MAK); Yashima (Nakai, 1 June 1941, TI); Kida-gun, Mt. Goken-zan (Matsui, 17 Aug. 1910, MAK). TOKUSHIMA. Naka-gun, Washiki-machi, Nakagawa (Yamazaki, 13 Aug. 1976, TI); Naka-gun, Hinotani-mura (Yoshinaga s.n., TI); Mima-gun, Handa, Okuyama (Kasai, 21 Sept. 1913, MAK); Oe-gun, Koyadaira-mura (Nikai 1295, TI). EHIME. Uma-gun, Oku-no-in (Araki 5743, SHO); Mt. Ishidzuchi-san (Makino, 22 July 1905, MAK); Nishiuwa-gun, Misaki-cho, Takaura, alt. 10m (Fukuoka 8691, KYO). KOUCHI. Kouchi, Engyo-ji (Shibuya, 28 July 1958, TI); Nangoku-shi, Shiraki-dani, alt. ca. 100m (Ohashi, 20 June 1973, TUSG); Tosa-gun, Kagamimura, Sadanaga (Kikuchi et al., 16 Oct. 1968, TUSG); Takaoka-gun, Mt. Yokokura-yama (Kimura, 29 Oct. 1939, TI); Ochi (Watanabe, 16 Sept. 1889, GH); Takaoka-gun, Niyodmura, Mori (Fujita 341, KYO); Hata-gun, Isl. Oki-no-shima (Yamawaki, 14 Oct. 1944, TI).

KYUSHU. OITA. Shimoge-gun, Masaka-mura (Yamamoto, 3 Aug. 1921, KYO); Mt. Hiko-san (Honda 1055, TI); near Beppu (Ishidoya, Aug. 1917, A). NAGASAKI. (Maximowicz in 1863, A; Faurie 3069, A); in insul. Iwosima leitum (Siebold s.n., L 908.118-1370-Syntype of L. Buergeri Miq.); Oomura-shi, Mt. Tara-dake, Housen-ji, alt. 650m (Shimada, 30 July 1958, KYO); Kokuzou (Higashisonogi-gun) (Makino, 23 Oct. 1904, MAK); Isl. Hiratojima (Hashimoto, 16-19 Aug. 1951, TI). MIYAZAKI. Takachiho (Momiyama, 8 Oct. 1958, TI); Nishiusuki-gun, Hinokage-machi, near Furuzono, alt. 150-250 m (Tateishi 3358, TI); Higashiusuki-gun, Toogoo-machi, Yamage (Murata 8590, TI); Koyu-gun, Kishiro-mura, at the foot of Mt. Osuzu-yama (Hurusawa 32, TI); Mt. Osuzu, alt. 900 m (Sako, 20 Oct. 1960, MAK), KUMAMOTO. Uto, Mt. Misumi-dake (Kôzuma, 23 June 1912, TNS); Kuma-gun, Itsuki-mura, Touji (Ohba & Akiyama 1031, TI). KAGOSIIIMA. Mt. Kirishima, alt. 100-1000 m (Tashiro in 1917, A); Mt. Takakuma-yama (Yamazaki, 12 Aug. 1942, TI); Kimotsuki-gun, Nejime-cho, Mt. Nokubidake, alt. 300m (Sako 2452, KYO); Kaseda-shi, Mt. Isoma-yama (Ohba & Akiyama 875, TI).

This species is greatly different from the other species of sect. Macrolespedeza. It is apparently a more woody plant than others and grows in shadier and/or rockier places. It blooms from (late June) July to September, earlier than others. One form with elliptic leaves with a round apex was named f. Oldhamii (Miq.) Sugimoto (=L. Oldhamii Miq.). The shape of leaves varies, however, so it is difficult to distinguish it based only on the shape of leaves.

Forma albiflora Honda in Bot. Mag. Tokyo 46: 421 (1932), ut "L. Buergeri Miq. var. Oldhamii (Miq.) Maxim. f. albiflora Honda"-Ohwi, Fl. Jap., 678 (1953); Fl. Jap. rev. ed., 789 (1965); Fl. Jap. Eng. ed., 559 (1965); Fl. Jap. new ed., 789 (1975).

L. Buergeri Miq. var. albiflora Honda in Bot. Mag. Tokyo 46: 421 (1932).

L. Buergeri Miq. f. leucantha Sugimoto, New Keys Jap. Trees, 463 (1961), nom.superfl.

Japanese name. Shirobana-kihagi (Honda, 1932).

Corolla white.

Rarely distributed in the range of this species.

Specimens examined.

HONSHU SHIZUOKA. Minamizaki (prov. Izu) (Sugimoto 21869, TI-Holotype of L. Buergeri var. Oldhamii f. albiflora Honda); Ikawa (prov. Sagami) (Sugimoto 20435, TI-Holotype of L. Buergeri var. albiflora Honda).

Forma angustifolia Makino in Bot. Mag. Tokyo 20: 41 (1906)-Hatusima in Mem. Fac. Agr. Kagoshima Univ. 6: 6 (1967)-Ohwi, Fl. Jap. new ed., 789 (1975).

L. Buergeri Miq. var. Kinashii Ohwi in Journ. Jap. Bot. 26: 234 (1951); Fl. Jap., 678 (1953); Fl. Jap. rev. ed., 789 (1965); Fl. Jap. Eng. ed., 559 (1965).

Japanese name. Tachige-kihagi (Ohwi, 1951), Hosoba-kihagi (Makino, 1906).

Stem and branches with patent hairs.

Distr. Sporadic throughout the range of the species.

Specimens examined.

SHIKOKU. TOKUSHIMA. Kamo (prov. Awa), Yoshii-mura (Nishimoto, 15 Sept. 1904, TI-Holotype of L. Buergeri Miq. f. angustifolia Makino).

Among the specimens cited under L. Buergeri, those referable to f. angustifolia are marked with an asterisk.

This form has branches and inflorescence-axes with patent hairs. Some plants with branches and inflorescence-axes with ascending hairs are also found sporadically.